Sauropoda Marsh, 1878
Definition- (Saltasaurus loricatus <- Melanorosaurus
readi) (Yates, 2007)
Other definitions- (Cetiosaurus oxoniensis, Diplodocus
longus, Brachiosaurus altithorax, Titanosaurus indicus
<- Plateosaurus engelhardti) (modified from McIntosh, 1997)
(Vulcanodon karibaensis + Diplodocus longus) (modified
from Salgado et al., 1997)
(Saltasaurus loricatus <- Plateosaurus engelhardti)
(Wilson, 2005; modified from Wilson and Sereno, 1998)
(Saltasaurus loricatus <- Jingshanosaurus xinwaensis,
Mussaurus patagonicus) (Sereno, 2007)
= Cetiosauria Owen, 1859
= Diplodocia Tornier, 1913
Comments- While Cope (1883) used Opisthocoela as an alternative
name for Sauropoda, it is actually a misspelling of Owen's (1860)
Opisthocoelia. As Opisthocoelia was erected to include not only "Cetiosaurus"
medius and Pelorosaurus but also the theropod Streptospondylus,
it is a better equivalent of Saurischia than Sauropoda. Tornier (1913)
used Diplodocia as an alternative name for Sauropoda.
References- Owen, 1859. On the Classification and Geographical
Distribution of the Mammalia. John W. Parker and Son, West Strand.
London. 103 pp.
Owen, 1860. Palaeontology, or a Systematic Summary of Extinct Animals
and their Geological Relations. Second Edition. Adam and Charles Black,
Edinburgh. 463 pp.
Cope, 1883. On the characters of the skull in the Hadrosauridae.
Proceedings of the Philadelphia Academy of Natural Sciences. 35, 97-107.
Tornier, 1913. Reptilia. Handwörterbuch der naturwissenschaften. 8,
315-376.
Marsh, 1878. Principal characters of American Jurassic dinosaurs. Part
I. American Journal of Science and Arts. 16, 411-416.
McIntosh, 1997. Sauropoda. In Currie and Padian (eds.). Encyclopedia of
Dinsoaurs. Academic Press, San Diego. 654-658.
Salgado, Coria and Calvo, 1997. Evolution of titanosaurid sauropods. I:
Phylogenetic analysis based on the postcranial evidence. Ameghiniana.
34(1), 3-32.
Wilson and Sereno, 1998. Early evolution and higher-level phylogeny of
sauropod dinosaurs. Society of Vertebrate Paleontology Memoir 5.
Journal of Vertebrate Paleontology. 18(2 suppl), 68 pp.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Sereno, 2007. Basal Sauropodomorpha: Historical and recent phylogenetic
hypotheses, with comments on Ammosaurus major (Marsh, 1889).
Special Papers in Palaeontology. 77, 261-289.
Yates, 2007. Solving a dinosaurian puzzle: the identity of Aliwalia
rex Galton. Historical Biology. 19(1), 93-123.
Leonerasaurus
Pol, Garrido and Cerda, 2011
L. taquetrensis Pol, Garrdo and Cerda, 2011
Early Pliensbachian, Early Jurassic
Upper Member of the Las Leoneras Formation, Chubut, Argentina
Holotype- (MPEF-PV 1663) partial dentary, three teeth, incomplete
axis, incomplete third cervical vertebra (40.8 mm), incomplete fourth
cervical vertebra (47.1 mm), incomplete fifth cervical vertebra (50.8
mm), partial sixth cervical vertebra, partial seventh cervical
vertebra, partial eighth cervical vertebra, ninth cervical vertebra,
tenth cervical vertebra, incomplete first dorsal vertebra (40.1 mm),
incomplete second dorsal vertebra (38 mm), incomplete third dorsal
vertebra (41.8 mm), partial fourth dorsal vertebra (42.3 mm), fifth
dorsal vertebra, ?sixth dorsal vertebra, incomplete mid dorsal vertebra
(42.7 mm), incomplete mid dorsal vertebra, incomplete mid dorsal
vertebra, fragmentary posterior dorsal vertebrae, dorsal ribs,
incomplete first sacral vertebra (46.9 mm), incomplete fused second and
third sacral vertebra (43, 43 mm), partial fourth sacral vertebra,
incomplete scapula, incomplete humerus (202.8 mm), incomplete ilia,
pubic fragment, incomplete ischium, femoral fragment, metatarsal I (67
mm), partial metatarsal II, incomplete pedal ungual
Diagnosis- (after Pol et al., 2011) anterior teeth with low,
spoon-shaped crowns (SI = 1.3); dorsosacral rib attached to
preacetabular process of ilium (also in Lufengosaurus);
primordial sacral neural arches positioned on anterior half of centrum;
caudosacral rib directed anterolaterally; deltopectoral crest low and
medially deflected along distal half; flattened ischial shafts (also in
Anchisaurus).
Comments- Pol et al. (2011) added Leonerasaurus to
Yates' sauropodomorph matrix and recovered it as sister to Melanorosaurus+Sauropoda.
More recently, McPhee et al. (2015) recovered Leonerasaurus as
sauropod more derived than Antetonitrus and Lessemsaurus,
but more basal than other included sauropods.
References- Pol, Garrido and Cerda, 2011. A new sauropodomorph
dinosaur from the Early Jurassic of Patagonia and the origin and
evolution of the sauropod-type sacrum. PLoS ONE. 6(1), e14572.
McPhee, Bonnan, Yates, Neveling and Choiniere, 2015. A new basal
sauropod from the pre-Toarcian Jurassic of South Africa: Evidence of
niche-partitioning at the sauropodomorph-sauropod boundary? Scientific
Reports. 5, 13224.
Chinshakiangosaurus
"Damalosaurus" Zhao,
1983
= "Damalasaurus" Zhao, 1985
"D. laticostalis" Zhao, 1985
?= "Damalasaurus magnus" Zhao, 1985
Etymology- Named for Damala Mountain in Qamdo, with the species
referring to wide ribs (laticostalis) or large size (magnus).
Early Jurassic
Middle Daye Group, Daye, Qamdo County, Tibet, China
Material- (~15 m) specimen including dorsal rib
Comments- The species is listed as "magnus" in the text, but
"laticostalis" in the plate caption. Olshevsky (1991) listed
"laticostalis as the type, though I consider it probable only one
species is intended by Zhao since two are never present in faunal lists
together (Zhao and Cheng, 1985; Zhang and Li, 1997; Fang et al., 2006).
Fang et al. use "laticostalis" for the species, suggesting it is the
intended one since Zhao is a coauthor. Zhao (1985) includes a
photograph of a dorsal rib in situ, which the length estimate above is
taken from assuming proportions like Barapasaurus. Glut (1997)
incorrectly stated it is Middle Jurassic in age. Weishampel et al.
(2004) list this as an undescribed sauropod in their faunal list.
Relationships- Stated by Zhao (1983, 1985) to be a primitive
sauropod. Chure and McIntosh (1989) listed it as a cetiosaurid, while
Lambert (1990) and Olshevsky (1991) listed it as a brachiosaurid. Glut
(1997) only listed it as Sauropoda incertae sedis. Fang et al. (2006)
place it in Cetiosauridae, noting again it's a primitive sauropod. As
those authors placing it in Brachiosauridae have never seen the
specimen, and brachiosaurids are currently unknown from Asia or the
Early Jurassic, that assignment seems less likely. Since Cetiosauridae
is currently viewed as a grade of basal eusauropods, "Damalosaurus" is
here placed as Sauropoda incertae sedis.
References- Zhao, 1983. Phylogeny and evolutionary stages of
Dinosauria. Acta Palaeontologica Polonica. 28(1-2), 295-306.
Zhao, 1985. The Jurassic Reptilia. In Wang, Cheng and Wang (eds.). The
Jurassic System of China. Stratigraphy of China. 11, 286-289, 347,
plates 10 and 11.
Zhao and Cheng, 1985. The Qamdo-Simao Subregion. In Wang, Cheng and
Wang (eds.). The Jurassic System of China. Stratigraphy of China. 11,
174-179.
Chure and McIntosh, 1989. A Bibliography of the Dinosauria (Exclusive
of the Aves) 1677-1986. Museum of Western Colorado Paleontology Series
#1. 226 pp.
Lambert, 1990. The Dinosaur Data Book. New York: Avon Books. 320 pp.
Olshevsky, 1991. A Revision of the Parainfraclass Archosauria Cope,
1869, Excluding the Advanced Crocodylia. Mesozoic Meanderings. 2, 196
pp.
Glut, 1997. Dinosaurs - The Encyclopedia. McFarland Press, Jefferson,
NC. 1076 pp.
Zhang and Li, 1997. Mesozoic Dinosaur Localities in China and Their
Stratigraphy. In Wolberg, Sump and Rosenberg (eds.). Dinofest
International, Proceedings of a Symposium sponsered by Arizona State
University. A Publication of The Academy of Natural Sciences. 265-273.
Weishampel, Barrett, Coria, Le Loeuff, Xu, Zhao, Sahni, Gomani and
Noto, 2004. Dinosaur Distribution. in Weishampel, Dodson and Osmolska
(eds.). The Dinosauria: Second Edition. University of California Press.
517-606.
Fang, Zhang, Lu, Han, Zhao and Li, 2006. Collision between the Indian
Plate and the paleo-Asian late and the appearance of Asian dinosaurs.
Geological Bulletin of China. 25(7), 862-873.
"Yibinosaurus" Ouyang vide
anonymous, 2001
"Y. zhoui" Ouyang vide anonymous, 2001
Toarcian, Early Jurassic
Dongyuemiao Member of Ziliujing Formation, Sichuan, China
Material- (Chongqing Museum of Natural History coll.) specimen
including dorsal vertebrae
Comments- This name originally appeared in a guidebook, with the
taxon stated to be under study by Ouyang. Ouyang (2003) later mentioned
it in his thesis as a new genus, which he places in an eponymous
subfamily within Vulcanodontidae. As he also includes the more basal Gongxianosaurus
in Vulcanodontidae, Ouyang's conception of the family may be more of a
grade. Ouyang further notes the anterior dorsal neural spines are
transversely expanded, which he believes indicates a possible
relationship to taxa with bifurcated spines like Datousaurus.
As Ouyang states "Yibinosaurus" is from the same locality as Gongxianosaurus,
the dorsal vertebrae, 51 articulated caudal vertebrae, scapulae and
ilium mentioned by Luo and Wang (1999) as Gongxianosaurus sp.
nov. may be the "Yibinosaurus" material.
References- Luo and Wang, 1999. New discovery on dinosaur
fossils from Early Jurassic, Sichuan, China. Chinese Science Bulletin.
44(23), 2182-2188.
Anonymous, 2001. Dinosaur Fossils from Chongqing Natural History Museum.
Ouyang, 2003. Skeletal characteristics of Mamenchisaurus youngi
and the systematics of mamenchisuarids. PhD thesis. Chengdu University
of Technology. 176 pp.
Zizhongosaurus?
"huangshibanensis" Li vide Li, Zhang and Cai, 1999
Jurassic
Red Beds, Sichuan, China
Comments- This name has only appeared in the abstract (and faunal
list?) of Li et al. (1999), so is a nomen nudum. Whether it is properly
referred to Zizhongosaurus is thus uncertain.
Reference- Li, Zhang and Cai, 1999. The Characteristics of the
Composition of the Trace Elements in Jurassic Dinosaur Bones and Red
Beds in Sichuan Basin, Geological Publishing House, Beijing. 155 pp.
undescribed sauropod (Joyce, 2004)
Early Jurassic
Hanson Formation, Antarctica
Material- (Augustana College coll.) (~9 m) vertebrae, pelvis, limb
elements
Comments- Discovered in 2003, to be described by Hammer.
References- Joyce, 2004. Digging for dinosaurs in Antarctica:
Giant bones suggest icy continent had warmer past. NPR, 1-23-2004.
Gongxianosaurus
Kunmingosaurus
Zhao vide Dong, 1992
K. wudingensis Zhao vide Dong, 1992
= "Kunmingosaurus wudingi" Zhao, 1985
= "Kunmingosaurus utingensjs" Zhao, 1985
= "Kunmingosaurus utingi" Zhen, Li and Rao, 1986
= "Kunmingosaurus wudingensis" anonymous, 1990 vide Olshevsky, 1991
Etymology- Kunming is the capital city of Yunnan, while Wuding
is the county and basin the specimen was found in.
Hettangian, Early Jurassic
Zhangjiawa Member of the Lufeng Formation, Hoshaofang, Wuding County,
Yunnan, China
Syntypes- (IVPP coll.) (~11 m) incomplete skeleton including
dorsal vertebrae, eight proximal caudal vertebrae, four chevrons,
ilium, pubis, femur (~847 mm), tibia, fibula, astragalus, metatarsal I,
phalanx I-1, pedal ungual I, metatarsal II, phalanx II-1, pedal ungual
II, metatarsal III, phalanx III-1, phalanx III-2, pedal ungual III,
metatarsal IV, phalanx IV-1, metatarsal V, phalanx V-1
....(BNHM,PZGR 74) dentary (Young, 1966)
....(BNHM,PZGR 75) dentary (Young, 1966)
Referred- ?(FMNH CUP 2042) maxilla (Barrett, 1999)
?(IVPP coll.) tooth (Upchurch and Barrett, 2000)
Comments- The species name is spelled "wudingi" in the text, and
"utingensjs" in the figure caption, which as noted by Chure and
McIntosh (1989) are probably unintentional variants, and the latter no
doubt a mispelling of "utingensis".
The skeleton was discovered by Su in 1954, but the dentaries were only
discovered later in 1960 and described and illustrated by Young (1966)
as specimens of Lufengosaurus magnus. Dong (1992) referred them
to the same individual as the postcrania since they came from the same
quarry. While Dong (and later Olshevsky, 2000) was incorrect in stating
Zhao described the taxon in 1985, his own book contains a diagnosis,
type and illustration so would seem to validate the nomenclature
(contra Barrett, 1999). Indeed, Upchurch et al. (2004) attribute the
name to Dong, 1992. A photo of part of the mounted skeleton is in Zhao
(1985), contra Barrett (1999). Dong et al. (1990) published a
photograph of the entire skeleton, and several are available online as
well. Li et al. (2010) note Kunmingosaurus lacks dorsal
pleurocoels and has a poorly developed fourth trochanter.
Simmons (1965) described a maxilla (FMNH CUP 2042) from Ta Ti in the
same beds as Yunnanosaurus robustus, which is similar to Kunmingosaurus
in having a sauropodan lateral plate and serrated, spatulate teeth with
a lingual ridge. Barrett (1999) redescribed it as a sauropod maxilla,
though he did not refer it to any genus. Upchurch and Barrett (2000)
described a referred tooth.
Relationships- Stated by Zhao (1985) to be a primitive sauropod.
Chure and McIntosh (1989) and Lambert (1990) listed it as a
cetiosaurid, while Olshevsky (1991) listed it provisionally as a
barapasaurid and Dong (1992) as a shunosaurine camarasaurid. Upchurch
and Barrett (2000) discussed it as a 'vulcanodontid', which was a grade
of basal sauropods in their scheme, and described a referred tooth.
Upchurch (1995) noted there was no published evidence suggesting it was
a vulcanodontid, however. He later (1998) noted that Kunmingosaurus'
teeth were more plesiomorphic than Barapasaurus, Shunosaurus,
Patagosaurus and other taxa in having only a shallow lingual
concavity and being quite labiolingually compressed. Barrett (1999)
considered the dentaries to be sauropod based on the dorsoventrally
expanded symphysis and lateral plate. Fang et al. (2006) list it as a
camarasaurid.
When entered into a modified version of Wilson's (2002) analysis, Kunmingosaurus
emerges more derived than Chinshakiangosaurus, Antetonitrus,
Lessemsaurus and Blikanasaurus but less than eusauropods
(so is not a cetiosaurid, barapasaurid, shunosaurid or camarasaurid).
It is thus related to taxa such as Gongxianosaurus, Kotasaurus,
Spinophorosaurus, Tazoudasaurus and Vulcanodon, so may be
considered vulcanodont-grade. Note the results agree with Upchurch's
(1998) statements, though his dental characters were not used by
Wilson. Specifically, Kunmingosaurus is more derived than Chinshakiangosaurus
based on the D-shaped teeth and reduced fourth trochanter, but less
derived than Shunosaurus based on no crown-to-crown occlusion,
straight dorsal ilial margin and unreduced pedal phalanges.
References- Simmons, 1965. The non-therapsid reptiles of the
Lufeng Basin, Yunnan, China. Fieldiana, Geology. 15, 1-93.
Young, 1966. On a new locality of the Lufengosaurus of Yunnan.
Vertebrata PalAsiatica. 10(1), 64-67.
Zhao, 1985. The Jurassic Reptilia. In Wang, Cheng and Wang (eds.). The
Jurassic System of China. Stratigraphy of China. 11, 286-289, 347,
plates 10 and 11.
Zhen, Li and Rao, 1986. Dinosaur footprints of Jinnin, Yunnan. Memoirs
of the Beijing Natural History Museum. 33, 19pp.
Chure and McIntosh, 1989. A Bibliography of the Dinosauria (Exclusive
of the Aves) 1677-1986. Museum of Western Colorado Paleontology Series
#1. 226 pp.
Dong, Hasegawa and Azuma, 1990. The Age of Dinosaurs in Japan and
China. Fukui, Japan: Fukui Prefectural Museum. 65 pp.
Lambert, 1990. The Dinosaur Data Book. New York: Avon Books. 320 pp.
Olshevsky, 1991. A Revision of the Parainfraclass Archosauria Cope,
1869, Excluding the Advanced Crocodylia. Mesozoic Meanderings. 2, 196
pp.
Dong, 1992. Dinosaurian Faunas of China. China Ocean Press (Beijing).
192 pp.
Upchurch, 1995. The evolutionary history of sauropod dinosaurs.
Philosophical Transactions of the Royal Society of London B. 349,
365-390.
Upchurch, 1998. The phylogenetic relationships of sauropod dinosaurs.
Zoological Journal of the Linnean Society. 124, 43-103.
Barrett, 1999. A sauropod dinosaur from the Lower Lufeng Formation
(Lower Jurassic) of Yunnan Province, People's Republic of China.
Journal of Vertebrate Paleontology. 19(4), 785-787.
Martin-Rolland, 1999. Les sauropodes chinois. Revue Paléobiologie,
Genève. 18(1), 287-315.
Olshevsky, 2000. An Annotated Checklist of Dinosaur Species by
Continent. Mesozoic Meanderings. 3, 157 pp.
Upchurch and Barrett, 2000. The evolution of sauropod feeding
mechanisms. in Sues (ed.). Evolution of Herbivory in Terrestrial
Vertebrates: Perspectives from the Fossil Record. Cambridge Press.
79-123.
Upchurch, Barrett and Dodson, 2004. Sauropoda. in Weishampel, Dodson
and Osmolska (eds.). The Dinosauria: Second Edition. University of
California Press. 259-322.
Weishampel, Barrett, Coria, Le Loeuff, Xu, Zhao, Sahni, Gomani and
Noto, 2004. Dinosaur Distribution. in Weishampel, Dodson and Osmolska
(eds.). The Dinosauria: Second Edition. University of California Press.
517-606.
Fang, Zhang, Lu, Han, Zhao and Li, 2006. Collision between the Indian
Plate and the paleo-Asian late and the appearance of Asian dinosaurs.
Geological Bulletin of China. 25(7), 862-873.
Archaeodontosaurus
Buffetaut, 2005
A. descouensi Buffetaut, 2005
Bathonian, Middle Jurassic
Isalo IIIb Formation, Madagascar
Holotype- (MHNDPal 2003-396) partial dentary, several teeth
Diagnosis- (after Buffetaut, 2005) dentary which is markedly
deeper anteriorly than posteriorly; at least 16 alveoli containing
prosauropod-like lanceolate teeth with strong, well-defined denticles,
a distinct lingual median ridge, and a finely wrinkled enamel.
Comments- Based on adding the taxon to Yates' sauropodomorph
analysis, this is a sauropod less derived than Shunosaurus. A
crown which expands markedly mesiodistally compared to the root may
indicate it is more derived than Chinshakiangosaurus. However,
Flynn et al. (2010) noted the locality and the mosaic of basal and
derived features might indicate a relationship to the Malagasy basal
archosauromorph Azendohsaurus, but the latter seems to lack a
lingual ridge on its teeth and wrinkled enamel.
Reference- Buffetaut, 2005. A new sauropod dinosaur with
prosauropod-like teeth from the Middle Jurassic of Madagascar. Bulletin
de la Societé Géologique de France. 176(5), 467-473.
Flynn, Nesbitt, Parrish, Ranivoharimanana and Wyss, 2010. A new species
of Azendohsaurus (Diapsida: Archosauromorpha) from the Triassic
Isalo Group of southwestern Madagascar: Cranium and mandible.
Palaeontology. 53(3), 669-688.
Eshanosaurus
Xu, Zhao and Clark, 2001
E. deguchiianus Xu, Zhao and Clark, 2001
Hettangian, Early Jurassic
Lower Lufeng Formation, Yunnan, China
Holotype- (IVPP V11579) (mandible ~170 mm) dentary, partial
splenial
Comments- This taxon was originally identified as a
therizinosaur, despite its Early Jurassic age. It may be a
sauropodomorph, especially since it lived so much earlier than any
other avetheropod, and the fact the much later therizinosaur Falcarius
is nonetheless less derived in respect to serration size, dentary
curvature and the absence of a lateral dentary ridge. Most recently,
Barrett (2009) notes that basal sauropodomorphs lack the anteriorly
extensive well defined lateral dentary shelf, anteriorly increasing
tooth size, more than thirty dentary teeth, lingual cingulum on teeth,
apicobasally oriented lingual ridge on teeth, and tooth roots that
expand mesiodistally to equal crown width.
References- Zhao and Xu, 1998. The oldest coelurosaurian.
Nature. 394, 234-235.
Xu, Zhao and Clark, 2001. A new therizinosaur from the Lower Jurassic
lower Lufeng Formation of Yunnan, China. Journal of Vertebrate
Paleontology. 21(3), 477-483.
Barrett, 2009. The affinities of the enigmatic dinosaur Eshanosaurus
deguchiianus from the Early Jurassic of Yunnan Province, People's
Republic of China. Palaeontology. 52(4), 681-688.
"Rutellum" Lhuyd, 1699
"R. impicatum" Lhuyd, 1699
Jurassic
Coral Rag, England
Material- (University of Oxford coll. #1352; lost) tooth
Comments- Lhuyd (1699) illustrated this as a fish tooth, but the
name is invalid as it was published before the earliest date for
Linnaean nomenclature. Gunther (1945) referred it to Cetiosaurus.
Delair and Sarjeant (2002) called it a cetiosaur. While the broad crown
excludes it from Diplodocoidea and Lithostrotia, further resolution is
difficult given the single figure. A crown which expands markedly
mesiodistally compared to the root may indicate it is more derived than
Chinshakiangosaurus.
References- Lhuyd, 1699. Lithophylacii Britannici Ichnographia,
sive lapidium aliorumque fossilium Britannicorum singulari figura
insignium. Gleditsch & Weidmann, London. 156 pp.
Gunther, 1945. Early Science in Oxford. Life and Letters of Edward
Lhuyd, vol. 14. the Author, Oxford.
Delair and Sarjeant, 2002. The earliest discoveries of dinosaurs; the
records re-examined. Proceedings of the Geologists' Association. 113,
185-197.
Amygdalodon
Cabrera, 1947
A. patagonicus Cabrera, 1947
Late Toarcian-Early Aalenian, Early-Middle Jurassic
Cerro Carnerero Formation, Chubut, Argentina
Lectotype- (MLP 46-VIII-21-1/2) posterior dorsal centrum (~150
mm)
Paralectotypes- (MLP 46-VIII-21-1/1) posterior dorsal centrum
(140 mm)
?(MLP 46-VIII-21-1/3) partial mid caudal vertebra (140 mm)
?(MLP 46-VIII-21-1/4) partial mid caudal vertebra (148 mm)
?(MLP 46-VIII-21-1/5) dorsal rib fragment
?(MLP 46-VIII-21-1/6) anterior dorsal neural spine
?(MLP 46-VIII-21-1/7) posterior cervical prezygapophysis
?(MLP 46-VIII-21-1/8) anterior or mid cervical centrum (258 mm)
?(MLP 46-VIII-21-1/9) dorsal rib fragment
?(MLP 46-VIII-21-1/10) flat fragment
?(MLP 46-VIII-21-1/11) dorsal rib fragment
?(MLP 46-VIII-21-1/12) maxillary tooth (22.7 mm)
?(MLP 46-VIII-21-1/13) dentary tooth
?(MLP 46-VIII-21-1/14) tooth root
?(MLP 46-VIII-21-1/15) upper tooth (23.8 mm)
?(MLP 46-VIII-21-1/16) tooth root
?(MLP 46-VIII-21-1/17) tooth
?(MLP 46-VIII-21-1/18; lost) tooth
?(MLP 46-VIII-21-1/19) incomplete pubis
?(MLP 46-VIII-21-1 coll.; three lost) three cervical rib fragments,
seven dorsal rib fragments
Referred- (MLP 36-XI-10-3/1) partial posterior dorsal vertebra
(~143 mm) (Casamiquela, 1963)
....(MLP 36-XI-10-3/3) dorsal rib fragment (Casamiquela, 1963)
?(MLP 36-XI-10-3/2) partial first sacral centrum (Casamiquela, 1963)
?(MLP 36-XI-10-3/4) distal tibia (178 mm wide) (Casamiquela, 1963)
Diagnosis- (after Rauhut, 2003) lateral walls of the neural
canal and centropostzygapophyseal laminae flared laterally posteriorly;
neural canal strongly flexed anteroposteriorly within the dorsal neural
arches.
(after Carballido and Pol, 2010) spoon-shaped teeth with low SI values
(1.34-1.49); enamel wrinkled forming a pattern of pits and narrow
apicobasal sulci; absence of denticles in both mesial and distal
margins (also in Kotasaurus and Shunosaurus); wear
facets extending mostly along one margin of the crowns.
Other diagnoses- Cabrera (1947) listed several characters in his
diagnosis. The teeth were said to be similar to Apatosaurus (as
Brontosaurus), but comparatively larger. However, the supposed Apatosaurus
tooth cited by Cabrera is from Camarasaurus, so was not
correctly associated with the Apatosaurus postcrania. In
addition, which postcrania the Amygdalodon teeth belonged to is
unknown. Apneumatic dorsal centra with pleurofossae are primitive. A
pubis "similar to atlantosaurids" is vague, but may indicate the
straight anterior margin noted in the text to be more like Apatosaurus
than Cetiosauriscus (as Cetiosaurus leedsi) or Camarasaurus.
A straight pubic shaft is also found in taxa such as Tazoudasaurus
however.
Comments- Piatnitzky discovered the referred material in 1936
and mentioned it that year, though it was not described until
Casamiquela did so in 1963. The type material was collected in 1936 and
described the following year by Cabrera as a new taxon of cetiosaurid.
Cabrera (1947) identified the dorsal neural spine as a caudal neural
spine, while Casamiquela (1963) believed it was a dorsal diapophysis.
The flat fragment has been identified as part of a scapula (Cabrera,
1947) and dorsal rib (Casamiquela, 1963), but Rauhut (2003) felt it
didn't match any sauropod element and placed it as Reptilia indet..
Camasiquela identified the sacral centrum as a caudal centrum. The
material was found in one area, but belongs to at least three
individuals, with the cervical prezygapophysis largest, the dorsals
medium sized, and the sacral smallest. Which individuals the other
material belongs to is unknown, though there are at least two sizes of
dorsal ribs. Whether all of the material belongs to the same taxon is
unknown, though it is all congruent with a basal sauropod.
Carballido and Pol (2010) recently found this taxon to be a sauropod
more derived than Chinshakiangosaurus but outside Gravisauria
based on a version of Wilson's analysis. Lang's (2008) unpublished
thesis finds it to be a shunosaurid though.
References- Piatnitzky, 1936. Estudio geológico de la region del
Río Chubut y del Río Genua. Boletin de Informaciones Petroleras.
13(137), 83-118.
Cabrera, 1947. Un saurópodo nuevo del Jurásico de Patagonia. Notas del
Museo de La Plata, Paleontología. 12, 1-17.
Casamiquela, 1963. Consideraciones acerca de Amygdalodon
Cabrera (Sauropoda. Cetiosauridae) del Jurásico Medio de la Patagonia.
Ameghiniana. 3(3), 79-95.
Rauhut, 2003. Revision of Amygdalodon patagonicus Cabrera, 1947
(Dinosauria, Sauropoda). Mitteilungen aus dem Museum fuer Naturkunde in
Berlin: Geowissenschaftliche Reihe. 6, 173-181.
Lang, 2008. Les cetiosaures (Dinosauria, Sauropoda) et les sauropodes
du Jurassique moyen: Revision systematique, nouvelles decouvertes et
implications phylogenetiques. Thesis. Museum National d’Histoire
Naturelle. 638 pp.
Carballido and Pol, 2010. The dentition of Amygdalodon patagonicus
(Dinosauria: Sauropoda) and the dental evolution in basal sauropods.
Comptes Rendus Palevol. 9, 83-93.
"Pulanesaura"
McPhee, Bonnan, Yates, Neveling and Choiniere, 2015
"P. eocollum" McPhee, Bonnan, Yates, Neveling and Choiniere, 2015
Hettangian-Sinemurian, Early Jurassic
Upper Elliot Formation, South Africa
Material- (at least two
individuals) (BP/1/6183) mid dorsal neural arch
..(BP/1/6183a) posterior dorsal neural arch
..(BP/1/6184) proximal ischium
..(BP/1/6186) pedal ungual I (120 mm)
..(BP/1/6191) metacarpal IV (82 mm)
?.(BP/1/6193) incomplete humerus
..(BP/1/6199) incomplete anterior or mid cervical vertebra
..(BP/1/6200) tibia (520 mm)
..(BP/1/6201) proximal caudal vertebra
..(BP/1/6202) ischium
..(BP/1/6204) tooth
..(BP/1/6205) chevron
..(BP/1/6207) tooth
..(BP/1/6210) incomplete ulna
..(BP/1/6646) proximal caudal vertebra (85 mm)
..(BP/1/6752) clavicle (193 mm)
..(BP/1/6768) dorsal rib
..(BP/1/6770) mid dorsal neural arch
..(BP/1/6882; intended holotype of "Pulanesaura eocollum") incomplete
anterior dorsal neural arch
..(BP/1/6980) tibia
..(BP/1/6983) proximal pedal ungual
..(BP/1/6984) incomplete anterior dorsal neural arch
..(BP/1/7366) incomplete ischium
..(BP/1/7741) mid or distal caudal vertebra
.. ilial fragment (lost), fibula (uncollected)
Diagnosis- (after McPhee et al., 2015) teeth with apicobasal
grooves on both labial and lingual surfaces teeth with denticles
restricted to apical third of crown; tooth crowns with extensive enamel
wrinkling easily discernible with naked eye; anterior or mid cervical
vertebra with anteroposteriorly short and dorsoventrally high neural
spines; anterior or mid cervical vertebra with dorsally raised and
obliquely set postzygapophyseal facets; anterior dorsal prezygapophyses
mediolaterally extensive and sheet-like; anterior dorsal anterior
infradiapophyseal fossae showing an externally constricted,
medially-tapering, pinched morphology; dorsoventrally tall anterior
dorsal neural spines, ~3 times taller than anteroposteriorly long;
middle to posterior dorsal neural arches with neural spines >1.6
times as high as long; proximal caudal vertebra with incipient
prezygadiapophyseal laminae; proximal caudal transverse processes
laterally restricted, triangular in shape and located on both neural
arch and centrum; proximal caudal centra almost twice as high
dorsoventrally as anteroposteriorly long; proximal caudal centra
lacking ventral sulcus; hyposphene on proximal caudal vertebra;
mediolaterally expansive radial fossa on proximal ulna; tibia with
proximal surface over twice as long anteroposteriorly than transversely
wide; tibia with similarly transversely restricted shaft; transversely
compressed first pedal ungual with convexly rounded proximoventral
margin.
Comments- Unfortunately, the description of "Pulanesaura" has no
mention of ZooBank and as of July 5
2020 it lacks an entry on the ZooBank webite. Thus
according to ICZN Article 8.5.3 (an electronic work must "be
registered in the Official Register of Zoological Nomenclature
(ZooBank) (see Article 78.2.4) and contain evidence in the work itself
that such registration has occurred"), "Pulanesaura eocollum" McPhee et
al., 2015 is a nomen nudum that may never be technically
valid
as its journal is not published physically.
McPhee et al. (2015) recovered "Pulanesaura" as sister to
vulcanodontids+gravisaurians using a version of Yates' matrix.
Reference- McPhee, Bonnan, Yates, Neveling and Choiniere, 2015.
A new basal sauropod from the pre-Toarcian Jurassic of South Africa:
Evidence of niche-partitioning at the sauropodomorph-sauropod boundary?
Scientific Reports. 5, 13224.
Gravisauria Allain and Aquesbi, 2008
Definition- (Tazoudasaurus naimi + Saltasaurus loricatus)
(modified from Allain and Aquesbi, 2008)
References- Allain and Aquesbi, 2008. Anatomy and phylogenetic
relationships of Tazoudasaurus naimi (Dinosauria, Sauropoda)
from the late Early Jurassic of Morocco. Geodiversitas. 30(2), 345-424.
Kotasaurus
Tazoudasaurus
Vulcanodontidae Cooper, 1984
Definition- (Vulcanodon karibaensis <- Diplodocus
longus) (modified from Allain et al., 2004)
References- Cooper, 1984. A reassessment of Vulcanodon
karibaensis Raath (Dinosauria: Saurischia) and the origin of the
Sauropoda. Palaeontologia Africana. 25, 203-231.
Allain, Aquesbi, Dejax, Meyer, Monbaron, Montenat, Rechir, Rochdy,
Russell and Taquet, 2004. A basal sauropod dinosaur from the Early
Jurassic of Morocco. Comptes Rendus Palevol. 3(3), 199-208.
Vulcanodon
Early Toarcian, Early Jurassic
Drakensburg Group, Zimbabwe
References- Yates, Hancox and Rubidge, 2004. First record of a
sauropod dinosaur from the Upper Elliot Formation (Early Jurassic) of
South Africa. South African Journal of Science. 100, 504-506.
Barapasauridae Halstead and Halstead, 1981
Reference- Halstead and Halstead, 1981. Dinosaurs. Blanford
Books. 170 pp.
Barapasaurus
Nebulasaurus Xing,
Miyashita, Currie, You, Zhang and Dong, 2015
= "Nebulasaurus" Xing, Miyashita, Currie, You, Zhang and Dong, 2013
online
N. taito Xing, Miyashita, Currie, You, Zhang and Dong,
2015
= "Nebulasaurus taito" Xing, Miyashita, Currie, You, Zhang and Dong,
2013 online
Aalenian-Bajocian, Middle Jurassic
Zhanghe Formation, Yunnan, China
Holotype- (LDRC-v.d.1) incomplete braincase
Diagnosis- (after Xing et al., 2015) exoccipitals nearly
excluding supraoccipital from foramen magnum (supraoccipital forms less
than a tenth of foramen magnum margin); supraoccipital not expanded
laterally between parietal and exoccipital.
Comments- The paper naming this taxon was originally available
online in 2013 but did not contain a ZooBank registration, so Nebulasaurus
taito was a nomen nudum until officially published in 2015 (ICZN
Article 8.5.3). Xing et al. (2015) added it to a version of Harris'
sauropod matrix and recovered Nebulasaurus as the sister to Spinophorosaurus,
in their phylogeny a non-neosauropod eusauropod.
Reference- Xing, Miyashita, Currie, You, Zhang and Dong, 2015. A
new basal eusauropod from the Middle Jurassic of Yunnan, China, and
faunal compositions and transitions of Asian sauropodomorph dinosaurs.
Acta Palaeontologica Polonica. 60(1), 145-154.
Spinophorosaurus
Gravisauria incertae sedis
Zizhongosaurus Dong, Zhou and Zhang, 1983
Z. chuanchengensis
Dong, Zhou and Zhang, 1983
Early Jurassic
Daanzhai Member of the Ziliujing Formation, Sichuan, China
Holotype- (IVPP V9067) posterior dorsal neural arch, partial
humerus, pubic fragment, fragments
Diagnosis- (after Dong et al., 1983) dorsal vertebrae with high
neural spine with an apex as an expanded plate that descends slightly
anteriorly and is concave posteriorly; lateral spine surface is
ornamented with dorsally radiating vertical striations; diapophyses are
well developed to form a right angle with the neural spine at the
neural arch; hyposphene present; relatively long forelimbs; humerus
with straight and rounded shaft.
Comments- Discovered in the 1980's. Originally referred to
Cetiosaurinae and more lately to Vulcanodontidae, Zizhongosaurus
has yet to be included in a phylogenetic analysis and will be difficult
to evaluate given the brief description and single figure.
References- Dong, Zhou and Zhang, 1983. The dinosaurian remains
from Sichuan Basin, China. Palaeontologia Sinica. 62(23), 1-145.
Eusauropoda Upchurch, 1995
= Bothrosauropodidae Janensch, 1929
= Bothrosauropodoidea Janensch, 1929 vide Kuhn, 1961
Definition- (Shunosaurus lii + Saltasaurus loricatus)
(Wilson, 2005; modified from Upchurch et al., 2004)
Other definitions- (Barapasaurus tagorei + Diplodocus
longus) (modified from Salgado et al., 1997)
(Saltasaurus loricatus <- Vulcanodon karibaensis)
(Sereno, 2007; modified from Wilson and Sereno, 1998)
Comments- Upchurch (1995) erected Eusauropoda for his clade
containing euhelopodids and neosauropods, but not Vulcanodon or
Barapasaurus. This makes the explicit inclusion of Barapasaurus
in Salgado et al.'s definition problematic. While both Upchurch et
al.'s and Wilson and Sereno's definitions have similar current content,
only the former equates to Eusauropoda in Upchurch's 1995 topology, so
is preferred here.
Janensch (1929) erected Bothrosauropodidae (which is invalid, as there
is no genus Bothrosauropus) for cetiosaurines, camarasaurines and
brachiosaurines.
References- Janensch, 1929. Material und Formegehalt der
Sauropoden in der Ausbeute der Tendaguru-Expedition, 1909-1912
[Material and figured content of sauropods in the yield of the
Tendaguru Expedition, 1909-1912]. Palaeontographica. Supplement VII
(1), 2(1), 3-34.
Kuhn, 1961. Die Familien der rezenten und fossilen Amphibien und
Reptilien. Verlaghus Meisenbach KG, Bamberg. 79 pp.
Upchurch, 1995. The evolutionary history of sauropod dinosaurs.
Philosophical Transactions of the Royal Society of London B. 349,
365-390.
Salgado, Coria and Calvo, 1997. Evolution of titanosaurid sauropods. I:
Phylogenetic analysis based on the postcranial evidence. Ameghiniana.
34(1), 3-32.
Wilson and Sereno, 1998. Early evolution and higher-level phylogeny of
sauropod dinosaurs. Society of Vertebrate Paleontology Memoir 5.
Journal of Vertebrate Paleontology. 18(2 suppl), 68 pp.
Upchurch, Barrett and Dodson, 2004. Sauropoda. in Weishampel, Dodson
and Osmolska (eds.). The Dinosauria (2nd edition). University of
California Press, Berkeley. 259-322.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Sereno, 2007. Basal Sauropodomorpha: Historical and recent phylogenetic
hypotheses, with comments on Ammosaurus major (Marsh, 1889).
Special Papers in Palaeontology. 77, 261-289.
Eusauropod indet. (Lapparent
and Zbyszewski, 1957)
Late Kimmeridgian, Late Jurassic?
Alcobaca Formation, Portugal
Material- (MMLT 602528;
syntype of Megalosaurus pombali)
proximal caudal vertebra (130 mm)
Comments- Lapparent and
Zbyszewski (1957) referred several vertebrae to their new species of Megalosaurus, M. pombali,
including "a very powerful posterior dorsal (Pl. XIII, fig. 31, 32, 33)
whose face has a somewhat less triangular shape (Torres Vedras
Museum)."
However, Mocho et al. (2017) instead referred this to "an anterior
caudal vertebra of an indeterminate sauropod."
References- Lapparent and Zbyszewski, 1957. Les dinosauriens du
Portugal. Mémoires des Services Géologiques du Portugal, nouvelle
série. 2, 1-63.
Mocho,
Royo-Torres, Escaso, Malafaia, de Miguel Chaves, Narváez, Pérez-García,
Pimentel, Silva and Ortega, 2017. Upper Jurassic sauropod record in the
Lusitanian Basin (Portugal): Geographical and lithostratigraphical
distribution.
Palaeontologia Electronica. 20.2.27A:
1-50.
Eusauropod
indet. (Lapparent and Zbyszewski, 1957)
Tithonian?, Late Jurassic?
Bombarral Formation?, Portugal
Material- (MG 4811; syntype of
Megalosaurus pombali)
mid-distal caudal vertebra (135 mm)
Comments- Lapparent and
Zbyszewski (1957) referred several vertebrae to their new species of Megalosaurus, M. pombali,
including "an amphicoelous and elongated posterior caudal (Pl. XIII,
fig. 30), very pinched ventrally, was recovered isolated at Albergaria.
It shows the powerful tail that this large megalosaur must have
had." However, Mocho et al. (2016) instead referred this to
Sauropoda indet., as "the presence of an anteroposteriorly short neural
arch seems to exclude this caudal vertebra from Theropoda, and its
general morphology more closely resembles that of sauropod middle and
posterior caudal vertebrae." They could differentiate it from Aragosaurus, Galveosaurus, Lusotitan, diplodocines and
titanosauriforms, but not Losillasaurus,
Lourinhasaurus, Turiasaurus or Zby.
Moch et al. state "In Albergaria dos Doze, the present railway
intersect Cretaceous sediments, “Cenomaniano inferior, Albiano,
Aptiano, Neocomiano”, nevertheless the referred point is close to the
river Arunca, which intersect the Upper Jurassic sediments of the
“Complexo Vale de Lagares” (Teixeira et al., 1968) that corresponds to
the Alcobaça Formation, Kimmeridgian to basal Tithonian in age."
Mocho et al. (2017) later stated it was "probably from the Bombarral
Formation."
References- Lapparent and Zbyszewski, 1957. Les dinosauriens du
Portugal. Mémoires des Services Géologiques du Portugal, nouvelle
série. 2, 1-63.
,
,
and ,
2016.
Systematic
review of Late Jurassic sauropods from the Museu Geológico collections
(Lisboa, Portugal).
Journal of Iberian Geology. 42:, 227-250.
Shunosauridae McIntosh, 1990 vide Durand, 2005
Shunosaurinae McIntosh, 1990
Reference- McIntosh, 1990. Sauropoda. In Weishampel, Dodson and
Osmolska (eds). The Dinosauria, Berkeley: University of California
Press. 345-401.
Durand, 2005. Major African contributions to Palaeozoic and Mesozoic
vertebrate palaeontology. Journal of African Earth Sciences. 43(1-3),
53-82.
Shunosaurus
Cardiodontidae Lydekker, 1895
Reference- Lydekker, 1895. On bones of a sauropodous dinosaur
from Madagascar. Quarterly Journal of the Geological Society of London.
51, 329-336.
Cardiodon
Rhoetosaurus
Longman, 1926
R. brownei Longman, 1926
Bajocian, Middle Jurassic
Hutton Sandstone Formation of the Injune Creek Group, Queensland,
Australia
Holotype- (QM F1659) incomplete ?axis (446 mm), cervical
vertebra, incomplete anterior dorsal vertebra (220 mm), three posterior
dorsal vertebrae (~180 mm), posterior dorsal centrum, partial posterior
dorsal centrum, two dorsal rib fragments, incomplete sacrum (16, 21
mm), fragmentary first caudal vertebra, fragmentary second caudal
vertebra, fragmentary third caudal vertebra, fragmentary fourth caudal
vertebra, incomplete fifth caudal vertebra (140 mm), sixth caudal
vertebra (136 mm), incomplete seventh caudal vertebra (137 mm),
incomplete eighth caudal vertebra (135 mm), ninth caudal centrum (135
mm), incomplete tenth caudal vertebra (140 mm), eleventh caudal
vertebra (142 mm), twelfth caudal centrum (160 mm), thirteenth caudal
centrum (150 mm), fourteenth caudal centrum (158 mm), fifteenth caudal
centrum (150 mm), sixteenth caudal vertebra (152 mm), seventeenth
caudal centrum (158 mm), eighteenth caudal centrum (157 mm), nineteenth
caudal centrum (155 mm), twentieth caudal centrum, distal caudal
vertebra (157 mm), five distal caudal vertebrae, fragmentary caudal
vertebrae, several chevron fragments, ilial fragments, incomplete pubes
(~1.2 m), ischial fragments, incomplete femur (~1.53 m), tibia, fibula,
astragalus, metatarsal I, phalanx I-1, pedal ungual I, metatarsal II,
phalanx II-1, pedal ungual II, metatarsal III, phalanx III-1, phalanx
III-2, phalanx III-3, pedal ungual III, metatarsal IV, phalanx IV-1,
pedal ungual IV, fragments
Diagnosis- (after Nair and Salisbury, 2009) multiple prominent
medial crests on the distal tibia; concentric proximal fossa on the
medial surface of the tibia; accessory fossae or grooves on the medial
face of pedal unguals.
Comments- The first remains were discovered in 1924, with
additional material in 1926 (Longman, 1927). Further material of the
type specimen was discovered starting in 1975 and was described with a
reanalysis of the taxon by Nair and Salisbury (2012). They find it to
be a non-neosauropod gravisaur.
References- Longman, 1926. A giant dinosaur from Durham Downs,
Queensland. Memoirs of the Queensland Museum. 8(3), 183-194.
Longman, 1927. The giant dinosaur: Rhoetosaurus brownei.
Memoirs of the Queensland Museum. 9(1), 1-18.
Longman, 1929. Palaeontological notes: Rhoetosaurus brownei.
Memoirs of the Queensland Museum. 9(3), 249.
Molnar, 1980. Australian late Mesozoic continental tetrapods: Some
implications. Mémoires de la Société Géologique de France. 139, 131-143.
Thulborn, 1985. Rhoetosaurus brownei the giant Queensland
dinosaur. in Vickers-Rich and van Tets (eds). Kadimakara: Extinct
Vertebrates of Australia. Pioneer Design Studio, Melbourne. 166-171.
Long, 1998. Dinosaurs of Australia and New Zealand and Other Animals of
the Mesozoic Era. UNSW Press. 192 pp.
Vickers-Rich, Rich and Hewitt, 1999. Wildlife of Gondwana. Indiana
University Press. 304 pp.
Nair and Salisbury, 2007. Osteology and biomechanics of the crus and
pes in Rhoetosaurus brownei Longman (Dinosauria: Sauropoda)
from the Middle Jurassic Injune Creek Group of Roma, south-western
Queensland. Geological Society of Australia Abstracts No. 85. 11th
Conference on Australian Vertebrate Evolution, Palaeontology and
Systematics. 58.
Nair and Salisbury, 2009. New anatomical information on Rhoetosaurus
brownei (Dinosauria: Eusauropoda), from the Jurassic Injune Creek
Group, near Roma, Queensland, Australia. Conference on Australian
Vertebrate Evolution, Palaeontology and Systematics. Geological Society
of Australia, Abstracts. 93, 76.
Nair and Salisbury, 2012. New anatomical information on Rhoetosaurus
brownei Longman, a gravisaurian sauropodomorph from the Middle
Jurassic of Queensland, Australia. Journal of Vertebrate Paleontology.
32(2), 369-394.
Bellusaurinae Dong, 1987
= "Bellusaurinae" Dong, 1986
Comments- Dong (1986) first used "Bellusaurinae" in a faunal
list. Though the official publication of Bellusaurinae is usually
attributed to Dong (1990), a shorter version was published earlier in
1987.
References- Dong, 1986. [unknown title and journal]
Dong, 1987. [untitled section on saurischians] in Zhao et al. (eds.).
Stratigraphy and Vertebrate Fossils of Xinjiang. Beijing, Institute of
Vertebrate Paleontology and Paleoanthropology. 28-30.
Dong, 1990. Sauropoda from the Kelamaili region of the Junggar Basin,
Xinjiang Autonomous Region. Vertebrata PalAsiatica. 28(1), 43-58.
Bellusaurus
Cetiosauridae Lydekker, 1888
Definition- (Cetiosaurus oxoniensis <- Saltasaurus
loricatus) (modified from Upchurch et al., 2004)
= Cetiosaurinae Lydekker, 1888 vide Janensch, 1929
References- Lydekker, 1888. Catalogue of the Fossil Reptilia and
Amphibia in the British Museum (Natural History). Part I. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamata,
Rhynchocephalia, and Proterosauria. British Museum (Natural History),
London. 309 pp.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute
der Tendaguru-Expedition, 1909-1912 [Material and figured content of
sauropods in the yield of the Tendaguru Expedition, 1909-1912].
Palaeontographica. Supplement VII (1), 2(1), 3-34.
Upchurch, Barrett and Dodson, 2004. Sauropoda. in Weishampel, Dodson
and Osmolska (eds.). The Dinosauria (2nd edition). University of
California Press, Berkeley. 259-322.
Cetiosaurus
Patagosaurus
Cetiosaurus? mogrebiensis
Chebsaurus
Lapparentosaurus
Ferganasaurus
"Lancangosaurus" Zhao,
1980
= "Lancanjiangosaurus" Zhao, 1983
= "Lancangjiangosaurus" Zhao, 1985
= "Lanchangjiangosaurus" Fang, Zhang, Lu, Han, Zhao and Li, 2006
"L. cachuensis" Zhao, 1985
Etymology- The genus references the Lancang Jiang, the local
name for the head of the Mekong River. The species name refers to the
Zaqu River, which is the upper reaches of the Mekong.
Middle Jurassic
Middle Dapuka Group, Dabuka, Qamdo County, Tibet, China
Material- (~15 m) specimen including skull, mandible, twelve teeth
and limb elements
specimen (Fang et al., 2006)
Comments- Though usually believed to be a preliminary name for Datousaurus
(e.g. Olshevsky, 1991), "Lancangosaurus" was first reported by Zhao
(1980) in relation to a Dapuka specimen, not the Xiashaximiao Datousaurus.
Listed characters are large skull, large spatulate teeth which decrease
gradually in size posteriorly, a thick mandible and robust limbs. Dong
et al. (1983) believed the teeth were congeneric with those from the
Wujiaba Quarry of the Shangshaximiao Formation, which they described as
belonging to Omeisaurus junghsiensis and O. fuxiensis.
Zhao (1985) figures teeth in situ, though the photograph is unclear.
Note Olshevsky (1991) switched the authorship for the variants
"Lancanjiangosaurus" and "Lancangjiangosaurus". Glut (1997) incorrectly
listed it as being Late Jurassic in age. Fang et al. (2006) note
additional material from lower in the same formation. Given the
etymology, either "Lancangosaurus" or "Lancangjiangosaurus" would be
the most accurate names.
Relationships- "Lancanjiangosaurus" was stated by Zhao (1983) to
be a bothrosauropodoid, in which he includes spatulate-toothed taxa
like camarasaurids and brachiosaurids. Zhao (1985) notes it is a
sauropod similar to Cetiosaurus. Chure and McIntosh (1989)
listed it as a cetiosaurid, as does Olshevsky (1991) provisionally,
while Lambert (1990) listed it as a brachiosaurid. Glut (1997) merely
referred it to Sauropoda incertae sedis. Most recently, Fang et al.
(2006) place it in Cetiosauridae. The known characters are found in
both basal eusauropods like Omeisaurus and camarasaurids,
though the age, locality and Zhao's opinion all favor the former
identification. Whether Dong et al. are correct in placing it close to Omeisaurus
is unknown, and for now I recommend keeping it as Eusauropoda incertae
sedis.
References- Zhao, 1980. [Mesozoic vertebrate-bearing beds and
stratigraphy of northern Xinjinag: Report of Paleontological Expedition
to Xinjiang IV.] Memoirs of the Institute of Vertebrate Palaeontology
and Palaeoanthropology, Academia Sinica A. 15, 1-119.
Dong, Zhou and Zhang, 1983. [Dinosaurs from the Jurassic of Sichuan].
Palaeontologica Sinica, New Series C. 162(23), 1-136.
Zhao, 1983. Phylogeny and evolutionary stages of Dinosauria. Acta
Palaeontologica Polonica. 28(1-2), 295-306.
Zhao, 1985. The Jurassic Reptilia. In Wang, Cheng and Wang (eds.). The
Jurassic System of China. Stratigraphy of China. 11, 286-289, 347,
plates 10 and 11.
Chure and McIntosh, 1989. A Bibliography of the Dinosauria (Exclusive
of the Aves) 1677-1986. Museum of Western Colorado Paleontology Series
#1. 226 pp.
Lambert, 1990. The Dinosaur Data Book. New York: Avon Books. 320 pp.
Olshevsky, 1991. A Revision of the Parainfraclass Archosauria Cope,
1869, Excluding the Advanced Crocodylia. Mesozoic Meanderings. 2, 196
pp.
Glut, 1997. Dinosaurs - The Encyclopedia. McFarland Press, Jefferson,
NC. 1076 pp.
Martin-Rolland, 1999. Les sauropodes chinois. Revue Paléobiologie,
Genève. 18(1), 287-315.
Fang, Zhang, Lu, Han, Zhao and Li, 2006. Collision between the Indian
Plate and the paleo-Asian late and the appearance of Asian dinosaurs.
Geological Bulletin of China. 25(7), 862-873.
Mamenchisauridae Young and Chao,
1972
= Omeisauridae Wilson, 2002
Definition- (Mamenchisaurus constructus <- Saltasaurus
loricatus) (Naish and Martill, 2007)
References- Young and Chao, 1972. [Mamenchisaurus hochuanensis
sp. nov.]. Institute of Vertebrate Palaeontology and Palaeoanthropology
Monographs, Series A. 8, 1-30.
Wilson, 2002. Sauropod dinosaur phylogeny: Critique and cladistic
analysis. Zoological Journal of the Linnean Society. 136, 217-276.
Naish and Martill, 2007. Dinosaurs of Great Britain and the role of the
Geological Society of London in their discovery: Basal Dinosauria and
Saurischia. Journal of the Geological Society, London. 164, 493-510.
Omeisaurus Young, 1939
O. jiaoi Jiang, Li, Peng
and Ye, 2011
Aalenian-Bajocian, Middle Jurassic
Xiashaximiao Formation, Sichuan, China
Holotype- (ZDM 5050) first dorsal vertebra (260 mm), second dorsal
vertebra (210 mm), third dorsal vertebra (220 mm), fourth dorsal
vertebra (210 mm), fifth dorsal vertebra (220 mm), sixth dorsal
vertebra (200 mm), seventh dorsal vertebra (210 mm), eighth dorsal
vertebra (207 mm), ninth dorsal vertebra (205 mm), tenth dorsal
vertebra (203 mm), eleventh dorsal vertebra ( 195mm), twelfth dorsal
vertebra, several dorsal ribs, five sacral vertebrae, first caudal
vertebra (110 mm), second caudal vertebra (93 mm), third caudal
vertebra (100 mm), fourth caudal vertebra (110 mm), fifth caudal
vertebra (115 mm), sixth caudal vertebra (115 mm), seventh caudal
vertebra (117 mm), eighth caudal vertebra (115 mm), ninth caudal
vertebra (120 mm), tenth caudal vertebra (120 mm), eleventh caudal
vertebra (115 mm), twelfth caudal vertebra (117 mm), thirteenth caudal
vertebra (123 mm), fourteenth caudal vertebra (120 mm), fifteenth
caudal vertebra (120 mm), sixteenth caudal vertebra (121 mm),
seventeenth caudal vertebra (128 mm), eighteenth caudal vertebra (128
mm), nineteenth caudal vertebra (124 mm), twentieth caudal vertebra
(120 mm), twenty-first caudal vertebra (120 mm), twenty-second caudal
vertebra (125 mm), twenty-third caudal vertebra (120 mm), twenty-fourth
caudal vertebra (120 mm), twenty-fifth caudal vertebra (115 mm),
twenty-sixth caudal vertebra (115 mm), twenty-seventh caudal vertebra,
twenty-eighth caudal vertebra , twenty-ninth caudal vertebra,
twenty-five chevrons, partial scapula (~1.62 m), coracoid, clavicle,
humeri (1.16, 1.12 m), radii (840, 820 mm), ulnae (~840, 875 mm), two
carpals, metacarpal I (190 mm), metacarpal II (250 mm), metacarpal III
(265 mm), metacarpal IV (225 mm), metacarpal V, ilium (~940 mm), pubis,
ischium (780 mm), femora (1.4 m), tibiae (875, 900 mm), fibulae (900,
935 mm), astragali, metatarsal I, metatarsal II, metatarsal III,
metatarsal IV, phalanges, unguals
Diagnosis- (after Jiang et al., 2011; note this is from the
English abstract, the Chinese actual diagnosis may include only a
portion of these) dorsal vertebrae tall and large; anterior dorsal
vertebrae (1 to 6) opisthocoelous; dorsal pleurocoels well developed
except for first; dorsal neural spines club-like; anterior dorsal
neural spines not bifurcated; caudal vertebrae relatively short and
thick; proximal caudal vertebrae slightly amphicoelous; first chevron
articulated to first caudal; first chevron shallow and small; first
caudal rib laterally directed, slightly fan-shaped; humerofemoral ratio
~.83; humerus long and straight, with slightly expanded proximal end
and greatly expanded distal end; deltopectoral crest well-developed and
low; ulnohumeral ratio ~.72; femur slender, with greatly expanded
proximal and distal ends; fourth trochanter well-developed;
tibiofemoral ratio ~.63.
Comments- This was discovered in 2000. The description has yet
to be translated to English and the taxon has not been analyzed
cladistically.
Reference- Jiang, Li, Peng and Ye, 2011. A new species of Omeisaurus
from the Middle Jurassic of Zigong, Sichuan. Vertebrata PalAsiatica.
49(2), 185-194.
Huangshanlong
Huang, You, Yang and Ren, 2014
H. anhuiensis Huang, You, Yang and Ren, 2014
Middle Jurassic
Hongqin Formation, Anhui, China
Holotype- (Anhui Paleontological Museum AGB5818) humerus (900 mm),
radius (520 mm), ulna (600 mm)
Diagnosis- (after Huang et al., 2014) transverse length of
proximal humerus 36% of humeral length; accessory processes near middle
of anterodistal humerus; radiohumeral ratio 58% of that of the humerus;
ulnohumeral ratio 67%; ulnar anteromedial process longer than
anterolateral process; ridges on anterior, anteromedial and
posterolateral faces of distal ulna.
Comments- The material was discovered in 2002. Huang et al.
(2014) described it as a new taxon belonging to Mamenchisauridae based
on several characters, but without a phylogenetic analysis.
Reference- Huang, You, Yang and Ren, 2014. A new sauropod
dinosaur from the Middle Jurassic of Huangshan, Anhui Province.
Vertebrata PalAsiatica. 52(4), 390-400.
Klamelisaurinae Zhao, 1993
Reference- Zhao, 1993. A new mid-Jurassic sauropod (Klamelisaurus
gobiensis gen. et sp. nov.) from Xinjiang, China. Vertebrata
PalAsiatica. 31(2), 132-138.
Klamelisaurus
Zhao, 1993
= "Klamelisaurus" SVP, 1990
K. gobiensis
Zhao, 1993
= "Klamelisaurus gobiensis" SVP, 1990
?= Kelmayisaurus "gigantus" Grady, 1993
Late Callovian-Early Oxfordian, Middle-Late Jurassic
Jiangjunmiao, Lower Shishugou Formation, Xinjiang, China
Holotype- (IVPP V9492) (~17 m,
adult) fragmentary teeth (lost), nine mid and posterior cervical
vertebrae, twelve dorsal vertebrae, three proximal dorsal ribs (two
lost), six sacral vertebrae, four partial proximal caudal vertebrae,
ten proximal caudal neural spines (five lost), eleven mid-distal caudal
centra (133, 147, ~156, 139, 116, 132, 135, 139, ~76, ~81 mm), four mid
to distal chevrons, incomplete scapulocoracoid, humerus (880 mm), ulna
(605 mm), radius (~540 mm), two ?carpals (lost), partial ilium,
incomplete pubis (lost), partial ischium, femora (one incomplete, one
distal; ~1.240 m), tibiae (one partial, one proximal; ~740 mm),
fibulae (one partial, one proximal; ~750 mm), partial astragalus,
calcaneum (lost), metatarsal I (~130 mm), metatarsal II/III (135 mm),
phalanx III-1(?) (63 mm), three pedal phalanges (~66, ~50, ~160 mm),
pedal ungual
Diagnosis- (after Moore et al.,
2020) scabrous, sheet-like anterior extensions of the SPRLs in
middle-posterior cervical vertebrae; oblong, blind pneumatic fossa just
anterior to the SDF in middle-posterior cervical vertebrae; triplicate
PCPLs in some posterior dorsal vertebrae; dorsally bifurcated SPDL in
middle and posterior dorsal vertebrae without confluence with either
the SPRL or the SPOL; shelf-like embayment of the proximomedial corner
of the anterior face of the humerus, bounded medially by a bulge;
flange-like ridge overhanging the medial surface of the shaft of
metatarsal I.
Comments- Discovered in 1982
and excavated in 1984. Olshevsky (1991) lists "Klamelisaurus
gobiensis" as a euhelopodid (in a sense including mamenchisaurs) "to be
described by Zhao C. C. from the Late Jurassic of the Dzungar Basin,
Xinjiang; known from a skeleton about 80% complete; SVP Bulletin
#149:45." Klamelisaurus
was officially described by Zhao (1993) as a brachiosaurid sensu lato
closest to camarasaurines but different enough to be in a monotypic
subfamily. Moore et al. (2020) redescribed Klamelisaurus
in depth, reporting some elements could not be located, others were
newly reported, the reported manual elements were pedal, and much of
the material was reconstructed making assessment of the original
anatomy impossible. Moore et al. recovered it as a euhelopodid
(including mamenchisaurs, almost always outside Neosauropoda) closest
to, and sometimes nested within, Mamenchisaurus.
Kelmayisaurus "gigantus"- Grady (1993) wrote about
visiting the Wucaiwan Formation of Xinjiang in
1987, stating "A few years earlier, Dale [Russell] noted in his field
book, a Kelmayisaurus gigantus,
a huge theropod measuring twenty-one meters from neck to tailtip (there
was no skull) had been found there." Olshevsky (DML, 1995)
reported
that "Dale says he's never heard of it", suggesting a misunderstanding
by Grady. A plausible solution was suggested by Greenfield
(online, 2021) that Russell's field book was referring to the sauropod Klamelisaurus gobiensis, whose
skull-less holotype was excavated from that formation in 1984.
The confusion in names could be due to the fact Klamelisaurus
was only officially published in April 1993, although the name
"Klamelisaurus gobiensis" had been unofficially proposed since at least
June 1990 (Olshevsky, 1991) so could have been in Russell's field book
and misinterpreted by Grady as a similar genus and species name.
Finally, while Klamelisaurus'
holotype was only estimated to be 17 meters by Zhao (1993) once
described, the anterior cervicals and distal caudals were not
preserved, so Russell's 21 meter report wouldn't be unreasonable
especially when done prior to the official description.
References- The Society of
Vertebrate Paleontology, 1990. Society of Vertebrate Paleontology News
Bulletin. Number 149 June 1990.
Olshevsky, 1991. A revision of the parainfraclass Archosauria Cope,
1869, excluding the advanced Crocodylia. Mesozoic Meanderings. 2, 196
pp.
Grady, 1993. The Dinosaur Project: The Story of the Greatest Dinosaur
Expedition Ever Mounted. Macfarlane Walter & Ross and The Ex Terra
Foundation. 261 pp.
Zhao, 1993. A new mid-Jurassic sauropod (Klamelisaurus gobiensis
gen. et sp. nov.) from Xinjiang, China. Vertebrata PalAsiatica. 31(2),
132-138.
Olshevsky, DML 1995. https://web.archive.org/web/20210524172844/http://dml.cmnh.org/1995Sep/msg00668.html
Moore, 2018. The systematics and evolution of Middle-Late Jurassic
sauropod dinosaurs from China, and macroevolutionary trends in
saurischian vertebral regionalization, shape, and pneumaticity. PhD
Thesis, The George Washington University. 538 pp.
Moore, Upchurch, Barrett, Clark and Xing, 2020. Osteology of Klamelisaurus gobiensis
(Dinosauria, Eusauropoda) and the evolutionary history of Middle-Late
Jurassic Chinese sauropods. Journal of Systematic Palaeontology.
18(16), 1299-1393.
Greenfield, online 2021. https://incertaesedisblog.wordpress.com/2021/02/11/what-is-kelmayisaurus-gigantus/
Qijianglong Xing,
Miyashita, Zhang, Li, Ye, Sekiya, Wang and Currie, 2015
Q. guokr Xing, Miyashita, Zhang, Li, Ye, Sekiya, Wang and
Currie, 2015
Late Jurassic
Suining Formation, Chongqing, China
Holotype- (QJGPM 1001) (subadult) partial skull, atlantal
intercentrum, axis (143.1 mm), third cervical vertebra (256 mm), fourth
cervical vertebra (230 mm), fifth cervical vertebra (310 mm), sixth
cervical vertebra (350 mm), seventh cervical vertebra (325 mm), eighth
cervical vertebra (305 mm), ninth cervical vertebra (330 mm), tenth
cervical vertebra (324 mm), eleventh cervical vertebra (394 mm),
twelfth cervical vertebra (345 mm), thirteenth cervical vertebra (373
mm), fourteenth cervical vertebra (348 mm), fifteenth cervical vertebra
(398 mm), sixteenth cervical vertebra (334 mm), seventeeth cervical
vertebra (312 mm), first dorsal vertebra, second dorsal vertebra, third
dorsal vertebra, fourth dorsal vertebra, fifth dorsal vertebra, sixth
dorsal vertebra, dorsal rib fragments, ~tenth caudal vertebra,
twenty-seven mid-distal (~15-41) caudal vertebrae, neural arch
fragments, chevron fragments, pubis, two pedal phalanges ?-1
Diagnosis- (after Xing et al., 2015) subequal anteroposterior
lengths of frontal and parietal (also in Omeisaurus and Shunosaurus);
parietal forming entire anterior margin of supratemporal fenestra (also
in Atlasaurus and Omeisaurus); absence of
frontoparietal fenestra and presence of postparietal foramen (also in Spinophorosaurus);
plate-like basipterygoid process oriented anteroventrally with
accessory tuber paralleling basal tuber; finger-like process lateral to
postzygapophyses in cervical vertebrae; pneumatopores in
spinodiapophyseal fossa in posterior cervical vertebrae; anterior
outline of neural spines of mid-caudal vertebra indented posteriorly
for more than half length of centrum (also in Mamenchisaurus);
pubis anteriorly concave such that distal end points more anteriorly
than ventrally.
Comments- The specimen was discovered in 2006. Xing et al.
(2016) recovered the taxon as a mamenchisaurid in a trichtomy with Chuanjiesaurus
and Mamenchisaurus+Yuanmousaurus.
Reference- Xing, Miyashita, Zhang, Li, Ye, Sekiya, Wang and
Currie, 2015. A new sauropod dinosaur from the Late Jurassic of China
and the diversity, distribution, and relationships of mamenchisaurids.
Journal of Vertebrate Paleontology. 35(1), e889701.
Mamenchisaurinae Young and Chao, 1972 vide McIntosh, 1990
Comments- McIntosh (1990) used Mamenchisaurinae in both of his
1990 works, but which was published first is uncertain.
References- Young and Chao, 1972. [Mamenchisaurus hochuanensis
sp. nov.]. Institute of Vertebrate Palaeontology and Palaeoanthropology
Monographs, Series A. 8, 1-30.
McIntosh, 1990. Species determination in sauropod dinosaurs with
tentative suggestions for their classification. In Carpenter and Currie
(eds.). Dinosaur Systematics: Perspectives and Approaches. Cambridge
University Press, Cambridge. 53-69.
McIntosh, 1990. Sauropoda. In Weishampel, Dodson and Osmolska (eds).
The Dinosauria, Berkeley: University of California Press. 345-401.
Mamenchisaurus
Tonganosaurus Li, Yang,
Liu and Wang, 2010
T. hei Li, Yang, Liu and Wang, 2010
Early Jurassic
Yimen Formation, Sichuan, China
Holotype- (MCDUT 14454) anterior axis, third cervical vertebra (168
mm), posterior fourth cervical centrum, posterior fifth cervical
vertebra, seventeenth cervical centrum (147 mm), second dorsal vertebra
(140 mm), fourth dorsal centrum (118 mm), eighth dorsal centrum (126
mm), ninth dorsal centrum (114 mm), tenth dorsal centrum (110 mm),
eleventh dorsal centrum (110 mm), partial first caudal vertebra (88
mm), second caudal vertebra (90 mm), third caudal vertebra (89 mm),
fourth caudal vertebra (90 mm), fifth caudal vertebra (97 mm), sixth
caudal vertebra (100 mm), seventh caudal vertebra (106 mm), partial
eighth caudal vertebra (110 mm), ten neural spine and rib fragments,
scapulae (one fragmentary; 760 mm), coracoid (208 mm), sternum (300
mm), humerus (628 mm), radius (498 mm), ulna (520 mm), ischia (570, 573
mm), femora (one partial; 840 mm), tibia (562 mm), fibula (566 mm),
metatarsal I (109 mm), pedal ungual I (170 mm), metatarsal II (166 mm),
metatarsal III (153 mm), metatarsal V (125 mm)
Diagnosis- (after Li et al., 2010) all presacrals pleurocoelous;
presacral bone texture solid, not cancellous; pleurocoels large and
deep; cervical centra slender; cervical centra ventrally keeled;
posterior cervical centra with well developed laminae and fossae;
dorsal vertebrae with well developed laminae and pleurocoels; cervical
and anterior dorsal centra opisthocoelous; mid dorsal centra
platycoelous; posterior dorsal and caudal centra amphicoelous;
forelimb/hindlimb ratio 0.80; humerus straight and robust;
deltopectoral crest well developed; femur straight and robust; fourth
trochanter well developed.
Comments- Found in 2007. Assigned by Li et al. to
Mamenchisauridae, specifically closest to Omeisaurus. However,
this has yet to be validated by a cladistic analysis.
Reference- Li, Yang, Liu and Wang, 2010. A new sauropod from the
Lower Jurassic of Huili, Sichuan, China. Vertebrata PalAsiatica.
48(3),185-202.
Xinjiangtitan
Wu, Zhou, Wings, Sekiya and Dong, 2013
X. shanshanensis Wu, Zhou, Wings, Sekiya and Dong, 2013
Bathonian-Oxfordian, Middle-Late Jurassic
Qigu Formation, Xinjiang, China
Holotype- (SSV12001) (~30-32 m) penultimate cervical vertebra (960
mm excl. condyle), last cervical vertebra (690 mm excl. condyle),
several partial cervical ribs, first dorsal vertebra (365 mm excl.
condyle), second dorsal vertebra (290 mm excl. condyle), third dorsal
vertebra (265 mm excl. condyle), fourth dorsal vertebra (255 mm excl.
condyle), (5th-8th combined 1.35 m excl. condyle) fifth dorsal
vertebra, sixth dorsal vertebra, seventh dorsal vertebra, eighth dorsal
vertebra, ninth dorsal vertebra (305 mm excl. condyle), tenth dorsal
vertebra (230 mm excl. condyle), eleventh dorsal vertebra (320 mm excl.
condyle), twelfth dorsal vertebra (340 mm excl. condyle), several
partial dorsal ribs, synsacrum (245, 260, 240, 270, 200 mm), first
caudal vertebra (210 mm), second caudal vertebra (220 mm), pubes,
proximal ischium, femur (1.65 m), tibia (80 mm), fibula, metatarsal IV
or V
Diagnosis- (after Wu et al., 2013) ventral median keel and
hypapophysis on penultimate cervical centrum; last two cervical
vertebrae highly elongated (combined 63% of femur+tibia); sacracostal
yoke developed except for first rib; robust femur (transverse width of
distal end 33% of length).
Comments- The holotype was discovered in 2012. When entered into
Harris' sauropod matrix, Xinjiangtitan emerged sister to Mamenchisaurus
(coded as a single OTU), with Omeisaurus less closely related.
Reference- Wu, Zhou, Wings, Sekiya and Dong, 2013. A new
gigantic sauropod dinosaur from the Middle Jurassic of Shanshan,
Xinjiang. Global Geology. 32(3), 437-446.
Tienshanosaurus
Young, 1937
T. chitaiensis Young, 1937
= Euhelopus chitaiensis (Young, 1937) Martin-Rolland, 1999
Late Oxfordian, Late Jurassic
Upper Shishugou Formation, Xinjiang, China
Holotype- (IVPP AS 40002-3) cervical vertebra (330 mm), cervical
central fragment, partial posterior cervical vertebra (~250 mm), two
cervical rib fragments, partial posterior dorsal vertebra, posterior
dorsal vertebra (120 mm), posterior dorsal vertebra (105 mm), dorsal
neural arch, several dorsal rib fragments, sacral fragment, partial
first caudal vertebra, second caudal vertebra, partial third caudal
vertebra, fourth caudal neural arch fragment, incomplete fifth caudal
vertebra (108 mm), sixth caudal vertebra (110 mm), seventh caudal
vertebra (105 mm), eighth caudal vertebra (90 mm), ninth caudal
vertebra (95 mm), tenth caudal vertebra (98 mm), incomplete eleventh
caudal vertebra (96 mm), twelfth caudal vertebra (93 mm), thirteenth
caudal vertebra (90 mm), fourteenth caudal vertebra (93 mm), incomplete
fifteenth caudal vertebra (94 mm), partial sixteenth caudal vertebra
(101 mm), partial twenty-eighth caudal vertebra, twenty-ninth caudal
vertebra (93 mm), thirtieth caudal vertebra (101 mm), thirty-first
caudal centrum (97 mm), thirty-second caudal vertebra (91 mm),
thirty-third caudal vertebra (89 mm), thirty-fourth caudal vertebra (89
mm), thirty-fifth caudal vertebra (90 mm), thirty-sixth caudal vertebra
(86 mm), thirty-seventh caudal central fragment, six caudal vertebrae,
caudal neural spines, two proximal chevrons, chevrons, incomplete
scapula (995 mm), coracoids (one partial, one incomplete), incomplete
humerus (680 mm), proximal radius, distal radius or ulna, incomplete
ilia (570, 565 mm), distal pubic fragment, ischia (one partial; 560
mm), proximal femur, distal ?femoral fragment, proximal tibia, distal
fibula
Referred- ?(IVPP V8301) seventh cervical vertebra, eighth
cervical vertebra, ninth cervical vertebra, tenth cervical vertebra,
cervical ribs, seventeen mid caudal vertebrae, chevrons (Dong, 1990)
Late Jurassic
Lamalishan, Xinjiang, China
?(IVPP V932) twentieth caudal vertebra (135 mm) (Young, 1958)
Diagnosis- (after Young, 1937) size moderate; scapula moderately
strong, weakly expanded distally; shaft subequal in length with the
expanded part plus coracoid; rather short forelimb; ilium with very
open anterior embayment; pelvic peduncle set forward; acetabulum
perforated but with a broad bony marginal area; ischium massive but not
distally expanded; fourth trochanter of femur weak or missing.
(after Dong, 1990) euhelopodid-style cervical vertebrae with long and
opisthocoelous centra, ventrally flat and lacking keels; neural spines
are low, or pseudospinus; large and long cervical ribs.
Comments- Discovered in 1930. Originally placed as a euhelopodid
or camarasaurid, based on a revised version of Wilson's analysis it may
be closer to basal taxa like Tazoudasaurus and Spinophorosaurus.
It certainly seems to differ from Euhelopus (e.g. in its
amphicoelous dorsals) and would have priority over the genus anyway,
contra Martin-Rolland (1999).
References- Young, 1937. A new dinosaurian from Sinkiang.
Palaeontologia Sinica. 105(2), 1-29.
Young, 1958. New sauropods from China. Vertebrata PalAsiatica. 2(1),
1-28.
Dong, 1990. Sauropoda from the Kelameili Region of the Junggar Basin,
Xinjiang Autonomous Region. Vertebrata PalAsiatica. 28(1), 43-58.
Martin-Rolland, 1999. Les sauropodes chinois. Revue Paléobiologie.
18(1), 287-315.
Losillasaurus
Turiasauria Royo-Torres, Cobos and
Alcalá, 2006
Definition- (Turiasaurus riodevensis <- Saltasaurus
loricatus) (Royo-Torres et al., 2006)
Reference- Royo-Torres, Cobos and Alcalá, 2006. A giant European
dinosaur and a new sauropod clade. Science. 314, 1925-1927.
Tendaguriidae Bonaparte, Heinrich and Wild, 2000
Reference- Bonaparte, Heinrich and Wild, 2000. Review of Janenschia
Wild, with the description of a new sauropod from the Tendaguru beds of
Tanzania and a discussion on the systematic value of procoelous caudal
vertebrae in the Sauropoda. Palaeontographica A. 256, 25-76.
Tendaguria
Bonaparte, Heinrich and Wild, 2000
T. tanzaniensis Bonaparte, Heinrich and Wild, 2000
Tithonian, Late Jurassic
Upper Dinosaur Member of the Tendaguru Formation, Tanzania
Holotype- (MB.R.2092.1, NB4) (~20 m) anterior dorsal vertebra
....(MB.R.2092.2, NB5) anterior dorsal vertebra
....(MB.R. coll.; lost) sacrum, ilium
Referred- ?(MB.R.2091.1-30, G1-G30) incomplete second caudal
vertebra (96.6 mm), incomplete third caudal vertebra (104 mm),
incomplete fourth caudal vertebra (110.7 mm), incomplete fifth caudal
vertebra (117.9 mm), incomplete sixth caudal vertebra (121 mm), partial
seventh caudal vertebra (129.2 mm), incomplete eighth caudal vertebra
(132.4 mm), incomplete ninth caudal vertebra (134.8 mm), tenth caudal
vertebra (137.3 mm), eleventh caudal vertebra (141.4 mm), twelfth
caudal vertebra (150.8 mm), thirteenth caudal vertebra (145 mm),
fourteenth caudal vertebra (148.4 mm), fifteenth caudal vertebra (143.4
mm), sixteenth caudal vertebra (144.5 mm), seventeeth caudal vertebra
(137.5 mm), eighteenth caudal vertebra (134.7 mm), nineteeth caudal
vertebra (133.5 mm), twentieth caudal vertebra (131.3 mm), twenty-first
caudal vertebra (~130.4 mm), twenty-second caudal vertebra (~133 mm),
twenty-third caudal vertebra (134.9 mm), twenty-fourth caudal vertebra,
twenty-fifth caudal vertebra (~140 mm), twenty-sixth caudal vertebra,
twenty-seventh caudal vertebra (~169 mm), twenty-eighth caudal vertebra
(~177.8 mm), twenty-ninth caudal vertebra (~179 mm), thirtieth caudal
vertebra, thirty-first caudal vertebra, two incomplete chevrons
(Janensch, 1929)
(MB.R.2091.31, G45) eighth or ninth cervical vertebra (Bonaparte,
Heinrich and Wild, 2000)
?(MB.R.2094, Oa12) sixth caudal vertebra (234 mm) (Janensch, 1929)
Diagnosis- (after Bonaparte et al., 2000) anterior dorsal
vertebrae opisthocoelous; neural spines low, continuous with
diapophysis and anteroposteriorly laminar; neural spines do not rise
above neural arch; diapophysis elongated with dorsal depression lateral
to prezygapophysis; diapophysis with deep anterior cavities and
shallower posterior cavities; well developed infradiapophyseal laminae;
epipophyses strong and thick, near dorsalmost border of vertebra and
connect with diapophyses.
Comments- Discovered between 1909 and 1912. First referred to Janenschia
robusta (as Gigantosaurus robustus) by Janensch (1929),
they may actually belong to that species. Bonaparte et al. (2000)
assigned it to a new neosauropod family, but preliminary analyses
suggest it may be related to near-neosauropod taxa like Jobaria
and Cetiosaurus instead. The caudal series is only tentatively
referred because it was associated with MB.R.2091.31.
References- Janensch, 1929. Material und Formegehalt der
Sauropoden in der Ausbeute der Tendaguru-Expedition, 1909-1912.
Palaeontographica. Supplement VII(1), 2(1), 3-34.
Bonaparte, Heinrich and Wild, 2000. Review of Janenschia Wild,
with the description of a new sauropod from the Tendaguru beds of
Tanzania and a discussion on the systematic value of procoelous caudal
vertebrae in the Sauropoda. Palaeontographica A. 256, 25-76.
Turiasaurus
Zby Mateus, Mannion and
Upchurch, 2014
Z. atlanticus Mateus, Mannion and Upchurch, 2014
Late Kimmeridgian, Late Jurassic
Amoreira-Porto Novo Member of Lourinha Formation, Portugal
Holotype- (ML 368) tooth (38x28x14 mm), mid-posterior cervical
neural arch fragment, incomplete proximal chevron, proximal scapula,
coracoid (569 mm), humerus (1.53 m), radius (1.04 m), ulna (1.06 m),
metacarpal I (365 mm), phalanx I-1 (82 mm), manual ungual I (172 mm),
phalanx ?II-1 (47 mm), metacarpal III (398 mm), metacarpal IV (382 mm)
Diagnosis- (after Mateus et al., 2014) posterior margin of
scapula just proximal to blade at an abrupt angle (~110 degrees) to
proximalmost posterior margin of scapular blade; humerus expands both
anteriorly and posteriorly at the distal half of the deltopectoral
crest, due to a prominent posterior bulge; laterodistal corner of
humerus forms an anteroposteriorly thin flange that does not expand as
far anteriorly as the rest of the distal end; manual ungual I
subrectangular in lateral view.
Comments- The holotype was discovered in 1996 and initially
referred to Turiasaurus riodevensis in an abstract (Mateus,
2009) before being described as a new taxon (Mateus et al., 2014).
However, the seven distinguishing characters listed by the latter
authors are only determinable in the holotypes of each species. As Turiasaurus
is only slightly later (Middle-Late Tithonian) and also from the
Iberian peninsula, it seems plausible the taxa are synonymous, though
this will require further comparable material to test.
Reference- Mateus, 2009. The sauropod dinosaur Turiasaurus
riodevensis in the Late Jurassic of Portugal. Journal of Vertebrate
Paleontology. 29(3), 144A.
Mateus, Mannion and Upchurch, 2014. Zby atlanticus, a new
turiasaurian sauropod (Dinosauria, Eusauropoda) from the Late Jurassic
of Portugal. Journal of Vertebrate Paleontology. 34(3), 618-634.
Neosauropoda Bonaparte, 1986
= Homalosauropodidae Janensch, 1929
= Homalosauropodoidea Janensch, 1929 vide Kuhn, 1961
Definition- (Diplodocus longus + Camarasaurus supremus)
(modified from Salgado et al., 1997)
Other definitions- (Diplodocus longus + Saltasaurus
loricatus) (Wilson, 2005; modified from Wilson and Sereno, 1998)
Comments- Janensch (1929) erected Homalosauropodidae (which is
invalid, as there is no genus Homalosauropus) for dicraeosaurines,
apatosaurines, diplodocines and titanosaurines.
References- Janensch, 1929. Material und Formegehalt der
Sauropoden in der Ausbeute der Tendaguru-Expedition, 1909-1912
[Material and figured content of sauropods in the yield of the
Tendaguru Expedition, 1909-1912]. Palaeontographica. Supplement VII
(1), 2(1), 3-34.
Kuhn, 1961. Die Familien der rezenten und fossilen Amphibien und
Reptilien. Verlaghus Meisenbach KG, Bamberg. 79 pp.
Bonaparte, 1986. Les Dinosaures (Carnosaures, Allosauridés, Sauropodes,
Cétiosauridés) du Jurassique moyen de Cerro Condor (Chubut, Argentine).
Annales de Paléontologie. 72, 247-289, 325-386.
Salgado, Coria and Calvo, 1997. Evolution of titanosaurid sauropods. I:
Phylogenetic analysis based on the postcranial evidence. Ameghiniana.
34(1), 3-32.
Wilson and Sereno, 1998. Early evolution and higher-level phylogeny of
sauropod dinosaurs. Society of Vertebrate Paleontology Memoir 5.
Journal of Vertebrate Paleontology. 18(2 suppl), 68 pp.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Haplocanthosauridae Bonaparte, 1999
Reference- Bonaparte, 1999. Evolución de las vértebras presacras
en Sauropodomorpha. Ameghiniana. 36(2), 115-187.
Haplocanthosaurus
Jobaria
Neosauropoda incertae sedis
Atlantosauridae Marsh, 1877
Atlantosaurinae Marsh, 1877 vide Steel, 1970
References- Marsh, 1877. Notice of new dinosaurian reptiles from
the Jurassic formation. American Journal of Science and Arts. 14,
514-516.
Steel, 1970. Part 14. Saurischia. Handbuch der
Paläoherpetologie/Encyclopedia of Paleoherpetology. Gustav Fischer
Verlag, Stuttgart. 87 pp.
Atlantosaurus
Bothriospondylidae Lydekker, 1895
Reference- Lydekker, 1895. On bones of a sauropodous dinosaur
from Madagascar. Quarterly Journal of the Geological Society of London.
51, 329-336.
Bothriospondylus
"Ischyrosaurus" Hulke,
1874 (preoccupied Cope, 1869)
"I." manseli Hulke vide Lydekker, 1888
= Ornithopsis manseli Lydekker 1888
= Pelorosaurus manseli (Lydekker, 1888) Lydekker, 1890
Early Kimmeridgian, Late Jurassic
Maple Ledge Shales of the Kimmeridge Clay Formation, England
Holotype- (NHMUK R41626) incomplete humerus (~770 mm)
Comments- Discovered in 1868. Note Cope (1869) unecessarily
renamed the plesiosaur Ischyrotherium Ischyrosaurus,
which predates Hulke's name. Barrett et al. (2010) determined it was
most likely a rebbachisaurid or basal titanosauriform so referred it to
Neosauropoda incertae sedis.
References- Cope, 1869. Synopsis of the extinct Batrachia and
Reptilia of North America. Transactions of the American Philosophical
Society. 14, 1-252.
Hulke, 1969. Note on a large saurian humerus from the Kimmeridge Clay
of the Dorset Coast. Quarterly Journal of the Geological Society. 25,
386-389.
Hulke, 1874. Note on a very large saurian limb-bone adapted for
progression on land, from the Kimmeridge Clay of Weymouth, Dorset.
Quarterly Journal of the Geological Society. 30, 16-17.
Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia in the
British Museum (Natural History). Part I. Containing the Orders
Ornithosauria, Crocodilia, Dinosauria, Squamata, Rhynchocephalia, and
Proterosauria. British Museum (Natural History), London. 309 pp.
Lydekker, 1890. Catalogue of the Fossil Reptilia and Amphibia in the
British Museum. Part IV. Containing the orders Anomodontia, Ecaudata,
Caudata and Labyrinthodontia; and Supplement. British Museum (Natural
History), London. 295 pp.
Barrett, Benson and Upchurch, 2010. Dinosaurs of Dorset: Part II, the
sauropod dinosaurs (Saurischia, Sauropoda) with additional comments on
the theropods. Proceedings of the Dorset Natural History and
Archaeological Society. 131, 113-126.
Ultrasaurus Kim,
1983
= "Ultrasaurus" Kim, 1981
U. tabriensis Kim, 1983
Aptian-Early Albian, Early Cretaceous
Gugyedong Formation, South Korea
Holotype- (DGBU-1973) proximal humerus (435 mm wide)
Referred- ?(DGBU coll.) caudal neural spine (Kim, 1983)
? rib fragments (Kim, 1988)
? cervical vertebra (Kim, 1988)
Comments- Discovered in 1977, initially identified as a sauropod
femur or tibia (Chang et al., 1982) or a proximal brachiosaurid ulna
(Kim, 1981, 1983). Paul (1988) notes it is actually a proximal humerus.
All modern authors have considered it an indeterminate sauropod, though
as Barrett et al. (2002) note, the absence of a well-developed
proximolateral humeral process excludes it from Somphospondyli. The
Cretaceous age may indicate it is a neosauropod.
References- Kim, 1981. Cretaceous dinosaur fossils discovered
from two dinosaur sites of Korea. Journal of the Geological Society of
Korea. 17, 297.
Chang, Seo and Park, 1982. Occurrence of a dinosaur limb bone near
Tabri, southern Korea. Journal of the Geological Society of Korea. 18,
195-202.
Kim, 1983. Cretaceous dinosaurs from South Korea. Journal of the
Geological Society of Korea. 19(3), 115-126.
Kim, 1988. Excavations and studies of dinosaur skeletons of Korea.
Abstracts at the Annual Meeting of the Paleontological Society of
Korea. 4(2), 168-169.
Paul, 1988. The brachiosaur giants of the Morrison and Tendaguru with a
description of a new subgenus, Giraffatitan, and a comparison
of the world's largest dinosaurs. Hunteria. 2, 1-14.
Lee, 1997. Reassessment of Ultrasaurus tabriensis, Kim 1983 and
the significance of Korean Sauropoda. Abstracts at the Annual Meeting
of the Paleontological Society of Korea. pg 14.
Lee, Yang and Park, 1997. Sauropod dinosaur remains from the Gyeongsang
Supergroup, Korea. Paleontological Society of Korea, Special
Publication. 2, 103-114.
Barrett, Hasegawa, Manabe, Isaji and Matsuoka, 2002. Sauropod dinosaurs
from the Lower Cretaceous of eastern Asia: Taxonomic and biogeographic
implications. Palaeontology. 45, 1197-1217.
Neosauropoda indet. (Hu,
1964)
Turonian, Late Cretaceous
Tashuikou, Ulanhsuhi Formation, Inner Mongolia, China
Material- (IVPP V2884.X; paratype of Chilantaisaurus tashuikouensis)
partial mid caudal centrum
Comments- Hu (1964) collected
this in 1960, apparently associated with Chilantaisaurus.
He said it was a carnosaurian middle caudal vertebra and
"most probably belong to the same species." Noting only the
subequal height and width (130 mm vs. 135 mm), his description was
surpassed by Benson and Xu (2008) who
stated it "has a stout and only weakly constricted centrum with
subrectangular articular surfaces, and the broken attachments of the
neural arch show that the latter was narrow." They said this
suggests "that the centrum belongs to a sauropod and should be
considered Sauropoda indet. (O.W.M.Rauhut, pers. comm.)". Indeed,
Rauhut (2003) had previously stated one of the vertebrae with Chilantaisaurus tashuikouensis'
specimen number "seems to represent a sauropod rather than a theropod."
References- Hu, 1964.
Carnosaurian remains from Alashan, Inner Mongolia. Vertebrata
PalAsiatica. 8, 42-63.
Rauhut, 2003. The interrelationships and evolution of basal theropod
dinosaurs. Special Papers in Palaeontology. 69, 213 pp.
Benson and Xu, 2008. The anatomy and systematic position of the
theropod dinosaur Chilantaisaurus tashuikouensis Hu, 1964 from
the Early Cretaceous of Alanshan, People’s Republic of China.
Geological Magazine. 145(6), 778-789.