Diplodocoidea Marsh, 1884 vide
Barrett and Upchurch, 1994
= "Diplodocoidea" Marsh, 1884 vide Upchurch, 1993
= Atlantosauria Haeckel, 1895
= Diplodocida Haeckel, 1895
Definition- (Diplodocus longus <- Saltasaurus
loricatus) (Wilson, 2005; modified from Wilson and Sereno, 1998)
Comments- Diplodocoidea was first used in Upchurch's (1993)
unpublished thesis, and later used in three papers in the same 1994 volume.
Upchurch (1994) uses it in quotes and notes it is only used as a label.
Barrett and Upchurch (1994) state its definition is in Upchurch's
thesis, but do not otherwise disclaim the name. Hunt et al. (1994) use
the superfamily later in the volume, but misspell it Diplodocoidae.
Upchurch (1995) is the usual reference given for the name.
References- Marsh, 1884. Principal characters of American
Jurassic dinosaurs. Part VII. Diplodocidae, a new family of the
Sauropoda. American Journal of Science. 27, 161-167.
Haeckel, 1895. Systematische Phylogenie der Wirbelthiere: (Vertebrata).
660 pp.
Upchurch, 1993. The anatomy, phylogeny and systematics of sauropod
dinosaurs. PhD thesis. University of Cambridge. 489 pp.
Hunt, Lockley, Lucas and Meyer, 1994. The global sauropod fossil
record. Gaia. 10, 261-279.
Barrett and Upchurch, 1994. Feeding mechanisms of Diplodocus.
Gaia. 10, 195-203.
Upchurch, 1994. Sauropod phylogeny and palaeoecology. Gaia. 10,
249-260.
Upchurch, 1995. The evolutionary history of sauropod dinosaurs.
Philosophical Transactions of the Royal Society of London, Series B.
349, 365-390.
Wilson and Sereno, 1998. Early evolution and higher-level phylogeny of
sauropod dinosaurs. Society of Vertebrate Paleontology Memoir 5.
Journal of Vertebrate Paleontology. 18(2 suppl), 68 pp.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Amazonsaurus
Amphicoeliidae Cope, 1877
Reference- Cope, 1877. On Amphicoelias, a genus of
saurians from the Dakota epoch of Colorado. Proceedings of the American
Philosophical Society. 17, 242-246.
Amphicoelias
Diplodocimorpha Calvo and
Salgado, 1995
Definition- (Limaysaurus tessonei + Diplodocus longus)
(modified from Calvo and Salgado, 1995)
Other definitions- (Rebbachisaurus garasbae + Diplodocus
longus) (modified from Taylor and Naish, 2005)
References- Calvo and Salgado, 1995. Rebbachisaurus tessonei
sp. nov. a new Sauropoda from the Albian-Cenomanian of Argentina; New
evidence on the origin of the Diplodocidae. GAIA. 11, 13-33.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea
(Dinosauria: Sauropoda). PaleoBios. 25(2), 1-7.
Rebbachisauridae Bonaparte,
1997
Definition- (Rebbachisaurus garasbae <- Diplodocus
longus) (Wilson, 2005; modified from Upchurch et al., 2004)
References- Bonaparte, 1997. Rayososaurus agrioensis
Bonaparte 1995. Ameghiniana. 34, 116.
Upchurch, Barrett and Dodson, 2004. Sauropoda. in Weishampel, Dodson
and Osmolska (eds.). The Dinosauria (2nd edition). University of
California Press, Berkeley. 259-322.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Rebbachisauroidea Bonaparte, 1997 vide Apesteguía, 2007
Definition- (Histriasaurus boscarollii + MACN PV N35 + Rebbachisaurus
garasbae + Limaysaurus tessonei) (after Apesteguía, 2007)
Comments- Apesteguía (2007) erected Rebbachisauroidea to contain
a node-based Rebbachisauridae and their basal relatives such as Histriasaurus
and MACN PV N35, but Rebbachisauridae is a stem-based clade which
includes these latter two taxa anyway. This would leave
Rebbachisauroidea as a subgroup of Rebbachisauridae with identical
known content. Since the term itself has limited utility, has not been
used since it was erected, and violates the ICZN by being inside
another superfamily (Diplodocoidea) and a family (Rebbachisauridae), it
is rejected here.
References- Bonaparte, 1997. Rayososaurus agrioensis
Bonaparte 1995. Ameghiniana. 34, 116.
Apesteguía, 2007. The sauropod diversity of the La Amarga Formation
(Barremian), Neuquén (Argentina). Gondwana Research. 12, 533-546.
Algoasaurus
Histriasaurus
Zapalasaurus
Nopcsaspondylus
Comahuesaurus
Carballido, Salgado, Pol, Canudo and Garrido, 2012
C. windhauseni Carballido, Salgado, Pol, Canudo and
Garrido, 2012
Aptian-Albian-Early Cretaceous
Puesto Quiroga Member of the Lohan Cura Formation, Neuquen, Argentina
Holotype- (MOZ-PV 6722) (at least three individuals) ~tenth-twelfth
dorsal neural arch
Paratypes- ..(MOZ-PV 6627) mid caudal vertebra (143 mm)
..(MOZ-PV 6628) mid caudal vertebra (165 mm)
..(MOZ-PV 6629) mid caudal vertebra
..(MOZ-PV 6630) proximal caudal vertebra (105 mm)
..(MOZ-PV 6631) partial proximal caudal vertebra (143 mm)
..(MOZ-PV 6632) mid caudal vertebra
..(MOZ-PV 6633) mid caudal vertebra (160 mm)
..(MOZ-PV 6634) mid caudal vertebra (166 mm)
..(MOZ-PV 6635) proximal caudal vertebra (90 mm)
..(MOZ-PV 6636) proximal caudal vertebra (127 mm)
..(MOZ-PV 6637) proximal caudal centrum (100 mm)
..(MOZ-PV 6638) mid caudal vertebra (168 mm)
..(MOZ-PV 6639) distal caudal vertebra (225 mm)
..(MOZ-PV 6640) proximal caudal centrum (107 mm)
..(MOZ-PV 6641) distal caudal vertebra (200 mm)
..(MOZ-PV 6642) mid caudal vertebra (210 mm)
..(MOZ-PV 6643) caudal vertebra
..(MOZ-PV 6644) caudal vertebra
..(MOZ-PV 6645) dorsal centrum
..(MOZ-PV 6646) mid caudal vertebra (198 mm)
..(MOZ-PV 6647) caudal vertebra
..(MOZ-PV 6648) proximal caudal vertebra (110 mm)
..(MOZ-PV 6649) mid caudal vertebra (187 mm)
..(MOZ-PV 6650) anterior dorsal vertebra (102 mm)
..(MOZ-PV 6651) dorsal centrum
..(MOZ-PV 6652) neural arch
..(MOZ-PV 6653) dorsal centrum (182 mm) or neural arch
..(MOZ-PV 6654) mid caudal vertebra (192 mm)
..(MOZ-PV 6657) mid-psoterior dorsal neural arch
..(MOZ-PV 6658) incomplete ischium
..(MOZ-PV 6659) pubic fragment
..(MOZ-PV 6660) pubic fragment
..(MOZ-PV 6661) fragmentary femur
..(MOZ-PV 6663) pubic fragment
..(MOZ-PV 6664) humeral fragment
..(MOZ-PV 6665) femur
..(MOZ-PV 6666) fragmentary femur
..(MOZ-PV 6667) pubic fragment
..(MOZ-PV 6669) two pubic fragments
..(MOZ-PV 6670) pubic fragment
..(MOZ-PV 6672) humeral fragment
..(MOZ-PV 6673) humeral fragment
..(MOZ-PV 6675) fragmentary ilium
..(MOZ-PV 6676) fragmentary ischium
..(MOZ-PV 6680) fragmentary ischium
..(MOZ-PV 6711) distal caudal vertebra (220 mm)
..(MOZ-PV 6712) humeral fragment
..(MOZ-PV 6713) partial ischium
..(MOZ-PV 6714) humeral fragment
..(MOZ-PV 6716) fragmentary ischium
..(MOZ-PV 6717) sternal plate
..(MOZ-PV 6719) partial ischium
..(MOZ-PV 6721) fragmentary femur
..(MOZ-PV 6723) humeral fragment
..(MOZ-PV 6727) partial fibula
..(MOZ-PV 6728) femur (1.13 m)
..(MOZ-PV 6729) mid caudal vertebra (180 mm)
..(MOZ-PV 6732) femur
..(MOZ-PV 6733) distal caudal vertebra (210 mm)
..(MOZ-PV 6734) distal caudal vertebra
..(MOZ-PV 6738) mid caudal vertebra (215 mm)
..(MOZ-PV 6740) proximal caudal vertebra
..(MOZ-PV 6741) incomplete first caudal vertebra (185 mm)
..(MOZ-PV 6743) pubis (805 mm)
..(MOZ-PV 6745) proximal caudal vertebra (90 mm)
..(MOZ-PV 6747) dorsal centrum
..(MOZ-PV 6748) proximal caudal vertebra
..(MOZ-PV 6751) dorsal centrum or chevron
..(MOZ-PV 6753) mid caudal vertebra (210 mm)
..(MOZ-PV 6755) femur
..(MOZ-PV 6756) partial posterior dorsal vertebra (183 mm)
..(MOZ-PV 6759) mid caudal vertebra (220 mm)
..(MOZ-PV 6760) proximal caudal vertebra (127 mm)
..(MOZ-PV 6761) femur
..(MOZ-PV 6762) humerus (675 mm)
..(MOZ-PV 6763) incomplete coracoid
..(MOZ-PV 6764) proximal tibia
..(MOZ-PV 6766) mid caudal vertebra (190 mm)
..(MOZ-PV 6767) proximal caudal vertebra (120 mm)
..(MOZ-PV 6778) fragmentary femur
Referred- ..(MOZ-PV 6718) tooth (Salgado et al., 2004)
Diagnosis- (after Carballido et al., 2012) anterior dorsal
centra with strong lateral constriction, resulting in a thin ventral
keel; anterior dorsal vertebrae with long prezygapophyses, in anterior
view covering around 3/4 of the transverse processes; anterior dorsal
vertebrae with two spinoprezygapophyseal laminae; anterior dorsal
vertebrae with two spinodiapophyseal laminae, an anterior and a
posterior one; anterior median lamina formed by three different
laminae, the anterior and posterior spinoprezygapophyseal laminae and
the anterior spinodiapophyseal; posterior dorsal centra with
centroprezygapophyseal lamina medially divided; posterior dorsal neural
arches with three spinopostzygapophyseal laminae; double contact
between posterior spinodiapophyseal lamina and lateral
spinopostzygapophyseal lamina; proximal caudal vertebrae with
well-developed prezygodiapophyseal fossa; caudal vertebrae with short
transverse process; robust humerus (robustness index 0.3); ischium with
straight shaft; shaft of ischium forming right angle with acetabulum;
ilial peduncle of ischium without constriction or neck.
Comments- The material was collected in 1995-1996 and 2002, and
initially described as Limaysaurus sp. by Salgado et al.
(2004). Carballido et al. (2012) described the rest of the material
once it was prepared, found that the characters used to refer it to Limaysaurus
were unknown in other rebbachisaurids. Therefore, they erected a new
genus for the material, and recovered Comahuesaurus as sister
to Khebbashia. Salgado et al. identified MOZ-PV 6741 as a last sacral
vertebra, but it was reidentified as the first caudal by Carballido et
al..
References- Salgado, Garrido, Cocca and Cocca, 2004. Lower
Cretaceous rebbachisaurid sauropods from the Cerro Aguada Leon (Lohan
Cura Formation), Neuquen Province, northwestern Patagonia, Argentina.
Journal of Vertebrate Palaeontology. 24(4), 903-912.
Carballido, Salgado, Pol, Canudo and Garrido, 2012. A new basal
rebbachisaurid (Sauropoda, Diplodocoidea) from the Early Cretaceous of
the Neuquen Basin; evolution and biogeography of the group. Historical
Biology. 24(6), 631-654.
Khebbashia Fanti, Cau,
Cantelli, Hassine and Auditore, 2015
Definition- (Limaysaurus tessonei + Nigersaurus
taqueti + Rebbachisaurus garasbae) (Fanti, Cau, Cantelli,
Hassine and Auditore, 2015)
Reference- Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New
information on Tataouinea hannibalis from the Early Cretaceous
of Tunisia and implications for the tempo and mode of rebbachisaurid
sauropod evolution. PLoS ONE. 10(4), e0123475.
Katepensaurus
Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado, 2013
K. goicoecheai Ibiricu, Casal, Martinez, Lamanna, Luna
and Salgado, 2013
Middle Cenomanian-Turonian, Late Cretaceous
Lower Member of Bajo Barreal Formation of the Chubut Group, Chubut,
Argentina
Holotype- (UNPSJB-PV 1007) incomplete frontal, two incomplete
anterior cervical vertebrae with fused ribs, incomplete mid cervical
vertebra with fused ribs, two cervical rib fragments, partial anterior
dorsal vertebra, partial anterior to mid dorsal vertebra, incomplete
anterior to mid dorsal neural arch, three incomplete mid to posterior
dorsal vertebrae (175 mm), four incomplete proximal caudal vertebrae,
proximal caudal neural arch, neural arch fragment, several partial
dorsal or proximal caudal neural spines, ?astragalar fragment,
?metapodial fragment, several fragments
Diagnosis- (after Ibiricu et al., 2013) internal lamina divides
lateral pneumatic fossa of mid to posterior dorsal centrum (also in Dinheirosaurus
and Supersaurus); vertical ridges on lateral surface of mid to
posterior dorsal vertebra, overlying neurocentral junction; pair of
laminae in mid to posterior dorsal parapophyseal centrodiapophyseal
fossa; mid to posterior dorsal transverse processes perforated by
elliptical fenestrae (that decrease in size posteriorly through dorsal
series and occupy approximately two-thirds of mediolateral process
width); well-defined, rounded fossae on lateral aspect of mid to
posterior dorsal postzygapophyses.
(after Ibiricu et al., 2015) posterior articular surface of mid to
posterior dorsal centrum with ventral portion wider than dorsal
portion, making it teardrop-shaped; anterior dorsal transverse
processes have laterodiapophyseal fossae homologous with more posterior
fenestrae.
Comments- Based on the quarry name, the holotype was discovered
in 2005. Ibiricu et al. (2013) referred Katepensaurus to
Rebbachisauridae and provisionally to Limaysaurinae. Ibiricu et al.
(2015) described further material of the holotype and added the taxon
to Whitlock's diplodocoid analysis to find it emerged as a
limaysaurine. However, Fanti et al. (2015) also used a version of that
analysis, and independently coding the originally described material,
recovered Katepensaurus as a basal rebbachisaurine. As this
latter analysis had the benefit of including Tataouinea and
information from Rebbachisaurus' redescription, which result is
better supported is uncertain without further work.
References- Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado,
2013. Katepensaurus goicoecheai gen. et sp. nov., a Late
Cretaceous rebbachisaurid (Sauropoda, Diplodocoidea) from central
Patagonia, Argentina. Journal of Vertebrate Paleontology. 33(6),
1351-1366.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications
for the tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE.
10(4), e0123475.
Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado, 2015. New material
of Katepensaurus goicoecheai (Sauropoda: Diplodocoidea) and its
significance for the morphology and evolution of Rebbachisauridae.
Ameghiniana. 52(4), 430-446.
Limaysaurinae Whitlock, 2011
Definition- (Limaysaurus tessonei <- Nigersaurus
taqueti) (modified from Whitlock, 2011
Reference- Whitlock, 2011. A phylogenetic analysis of
Diplodocoidea (Saurischia: Sauropoda). Zoological Journal of the
Linnean Society. 161(4), 872-915.
Limaysaurus
Cathartesaura
Rayososaurus
Rebbachisaurinae Bonaparte,
1997 vide Fanti, Cau, Cantelli, Hassine and Auditore, 2015
Definition- (Rebbachisaurus garasbae <- Limaysaurus
tessonei) (Fanti, Cau, Cantelli, Hassine and Auditore, 2015)
= Nigersaurinae Whitlock, 2011
Definition- (Nigersaurus taqueti <- Limaysaurus tessonei)
(modified from Whitlock, 2011
References- Bonaparte, 1997. Rayososaurus agrioensis
Bonaparte 1995. Ameghiniana. 34, 116.
Whitlock, 2011. A phylogenetic analysis of Diplodocoidea (Saurischia:
Sauropoda). Zoological Journal of the Linnean Society. 161(4), 872-915.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications
for the tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE.
10 (4), e0123475.
Nigersaurus
Demandasaurus
Torcida Fernández-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola
and Salgado, 2011
D. darwini Torcida Fernández-Baldor, Canudo, Huerta,
Montero, Pereda Suberbiola and Salgado, 2011
Late Barremian-Early Aptian, Early Cretaceous
Castrillo la Reina Formation, Burgos, Spain
Holotype- (MDS-RVII) (adult, ~10-12 m) premaxillae, dentary, six
teeth (~2.6x.6x?, ~1.5x.6x.4, 1.9x.6x.4, 1.6x.6x.4, 1.5x.6x.4, 1.2x.5x?
mm), axis (100 mm), mid cervical vertebra (270 mm), posterior cervical
vertebra, five cervical ribs, two posterior dorsal vertebrae (150, 150
mm), nine dorsal ribs, first caudal vertebra (145 mm), second caudal
vertebra (92 mm), six proximal caudal vertebrae (98, 100 mm), eleven
mid caudal vertebrae (150, 165, 165, 185, 175, 175 mm), nine chevrons
(315, 355, 370, 340, ~135, 205, 150, 131, 130 mm), ischia, femur (1.08
m)
Diagnosis- (after Torcida Fernandez-Baldor et al., 2011) teeth
ornamented with longitudinal crests on labial and lingual faces, and
bear mesial and distal carinae; posterior cervical vertebrae have
infraprezygapophyseal chamber with forked vertical accessory lamina;
posterior cervical vertebrae have rhombic accessory structure where
centroprezygapophyseal, prezygodiapophyseal and spinoprezygapophyseal
laminae are connected, dorsally to prezygapophyses; presacral
centroprezygapophyseal laminae divided; presence in mid dorsals of two
large neural arch pneumatic foramina that pass completely through
neural arch anteroposteriorly; presence of two large, deep pneumatic
cavities, divided by accessory laminae, in proximal caudal transverse
processes; proximal caudal anterior centroparapophyseal, posterior
centroparapophyseal and posterior centrodiapophyseal laminae very wide
and make contact posteriorly and ventrally with diapophysis; two
parallel laminae running in anteroposterior direction, an upper one
from prezygapophysis to base of centropostzygapophyseal, and lower one
from base of prezygapophysis to dorsal surface of proximal caudal
centra; two parallel crests running anteroposteriorly on lateral faces
of mid-distal caudal centra.
Comments- Found in 2002-2004. Torcida et al. (2011) added Demandasaurus
to Sereno et al.'s rebbachisaurid matrix and found it to be a
nigersaurine. This general relationship has been recovered in
subsequent analyses as well, though the redescription of Rebbachisaurus
has led to the subfamily being renamed Rebbachisaurinae. Most recently,
Fanti et al. (2015) found it to be sister to Rebbachisaurus+Tataouinea
using a version of Whitlock's diplodocoid analysis.
References- Pereda Suberbiola, Torcida, Izquierdo, Huerta,
Montero and Pérez, 2003. First rebbachisaurid dinosaur (Sauropoda,
Diplodocoidea) from the Early Cretaceous of Spain:
Palaeobiogeographical implications. Bulletin de la Société Géologique
de France. 174, 471-479.
Torcida Fernández-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola
and Salgado, 2011. Demandasaurus darwini, a new rebbachisaurid
sauropod from the Early Cretaceous of the Iberian Peninsula. Acta
Palaeontologica Polonica. 56(3), 535-552.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications
for the tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE.
10(4), e0123475.
Rebbachisaurus
Tataouinea Fanti,
Cau, Hassine and Contessi, 2013
T. hannibalis Fanti, Cau, Hassine and Contessi, 2013
Early Albian, Early Cretaceous
Oum ed Diab Member of the Ain el Guettar Formation, Tunisia
Holotype- (ONM DT 1-36) (adult) partial synsacrum (~800 mm),
partial first caudal vertebra, partial second caudal vertebra,
incomplete third caudal vertebra, incomplete fourth caudal vertebra,
fifth caudal vertebra, incomplete sixth caudal vertebra (140 mm),
incomplete seventh caudal vertebra (130 mm), eighth caudal vertebra
(130 mm), ninth caudal vertebra (190 mm), tenth caudal vertebra (160
mm), eleventh caudal vertebra (230 mm), twelfth caudal vertebra (230
mm), thirteenth caudal vertebra (240 mm), fourteenth caudal vertebra
(210 mm), fifteenth caudal vertebra (240 mm), sixteenth caudal vertebra
(250 mm), incomplete seventeenth caudal vertebra (220 mm), partial
ilia, proximal ischia
Diagnosis- (after Fanti et al., 2013) elliptical foramen in
lateral surface of fourth sacral neural spine penetrating its camerate
sector; proximalmost five caudal vertebrae with large elliptical
pleurocoel (modified after Fanti et al., 2015); 'lateral lamina' in
proximal caudal neural spines is 'inverted Y'-shaped, formed by
spinoprezygapophyseal and spinodiapophyseal laminae merging in ventral
third of spine and bordering a triangular fossa; ischium with large
elliptical foramen in medial surface of ilial peduncle.
Comments- The holotype was discovered in 2011, preliminarily
named and described in 2013, then described in depth two years later.
While Fanti et al. (2013) originally described what they thought were
the first five caudal vertebrae, they later (2015) identified the real
second caudal that belongs in the middle of that series, and eleven
subsequent caudals that weren't excavated at the time. Both papers used
a version of Whitlock's diplodocoid analysis to recover Tataouinea
in similar positions within Rebbachisauridae, though the proper
subfamily name changed from Nigersaurinae to Rebbachisaurinae based on Rebbachisaurus
itself being restudied and thus reclassified.
References- Fanti, Cau, Hassine and Contessi, 2013. A new
sauropod dinosaur from the Early Cretaceous of Tunisia with extreme
avian-like pneumatization. Nature Communications. 4, 2080.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications
for the tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE.
10(4), e0123475.
Flagellicaudata Harris and
Dodson, 2004
Definition- (Dicraeosaurus hansemanni + Diplodocus
longus) (Wilson, 2005; modified from Harris and Dodson, 2004)
Reference- Harris and Dodson, 2004. A new diplodocoid sauropod
dinosaur from the Upper Jurassic Morrison Formation of Montana, USA.
Acta Palaeontologica Polonica. 49(2), 197-210.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Dicraeosauridae Huene, 1927
Definition- (Dicraeosaurus hansemanni <- Diplodocus
longus) (Wilson, 2005; modified from Sereno, 1998)
References- Huene, 1927. Short review of the present knowledge
of the Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Sereno, 1998. A rationale for phylogenetic definitions, with
application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch
für Geologie und Paläontologie, Abhandlungen. 210(1), 41-83.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
undescribed dicraeosaurid (Gallina, Apesteguía, Haluza,
Canale and Otero, 2014)
Late Berriasian-Valanginian, Early Cretaceous
Bajada Colorada Formation, Neuquen, Argentina
Material- (MMCH-Pv coll.) incomplete skull, incomplete mandible,
vertebrae
Reference- Gallina, Apesteguía, Haluza, Canale and Otero, 2014.
The sauropod fauna of the Bajada Colorada Formation
(Berriasian-Valanginian), Neuquen Province, Argentina. Jornadas
Argentinas de Paleontologia de Vertebrados. Ameghiniana. 51(6)
suplemento, 11.
Dyslocosaurus
Suuwassea
Amargasaurus
Brachytrachelopan
Dicraeosaurinae Huene, 1927 vide Janensch, 1929
References- Huene, 1927. Short review of the present knowledge
of the Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute
der Tendaguru-Expedition, 1909-1912 [Material and figured content of
sauropods in the yield of the Tendaguru Expedition, 1909-1912].
Palaeontographica. Supplement VII(1) 2(1), 3-34.
Dicraeosaurus
Diplodocidae Marsh, 1884
Definition- (Diplodocus longus <- Dicraeosaurus
hansemanni) (Wilson, 2005; modified from Sereno, 1998)
= Apatosauridae Huene, 1927
References- Marsh, 1884. Principal characters of American
Jurassic dinosaurs. Part VII. Diplodocidae, a new family of the
Sauropoda. American Journal of Science. 27, 161-167.
Huene, 1927. Short review of the present knowledge of the Sauropoda.
Memoirs of the Queensland Museum. 9(1), 121-126.
Sereno, 1998. A rationale for phylogenetic definitions, with
application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch
für Geologie und Paläontologie, Abhandlungen. 210(1), 41-83.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry
Rogers and Wilson (eds.). The Sauropods: Evolution and Paleobiology.
University of California Press, Berkeley. 15-49.
Amphicoelias
"Probardodiplodocus"
Apatosaurinae Huene, 1927 vide
Janensch, 1929
Definition- (Apatosaurus ajax <- Diplodocus longus)
(modified from Taylor and Naish, 2005)
Other definitions- (Apatosaurus louisae <- Diplodocus
longus) (Wilson and Allain, 2015)
= Apatosaurinae sensu Wilson and Allain, 2015
Definition- (Apatosaurus louisae <- Diplodocus longus)
References- Huene, 1927. Short review of the present knowledge
of the Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute
der Tendaguru-Expedition, 1909-1912 [Material and figured content of
sauropods in the yield of the Tendaguru Expedition, 1909-1912].
Palaeontographica. Supplement VII(1) 2(1), 3-34.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea
(Dinosauria: Sauropoda). PaleoBios. 25(2), 1-7.
Wilson and Allain, 2015. Osteology of Rebbachisaurus garasbae
Lavocat, 1954, a diplodocoid (Dinosauria, Sauropoda) from the early
Late Cretaceous-aged Kem Kem beds of southeastern Morocco. Journal of
Vertebrate Paleontology. 35(4), e1000701.
DOI:10.1080/02724634.2014.1000701
Atlantosaurus
Apatosaurus
Brontosaurus
Diplodocinae Marsh, 1884 vide
Janensch, 1929
Definition- (Diplodocus longus <- Apatosaurus ajax)
(modified from Taylor and Naish, 2005)
Other definitions- (Diplodocus longus <- Apatosaurus
louisae) (Wilson and Allain, 2015)
= Diplodocinae sensu Wilson and Allain, 2015
Definition- (Diplodocus longus <- Apatosaurus louisae)
References- Marsh, 1884. Principal characters of American
Jurassic dinosaurs. Part VII. Diplodocidae, a new family of the
Sauropoda. American Journal of Science. 27, 161-167.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute
der Tendaguru-Expedition, 1909-1912 [Material and figured content of
sauropods in the yield of the Tendaguru Expedition, 1909-1912].
Palaeontographica. Supplement VII(1) 2(1), 3-34.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea
(Dinosauria: Sauropoda). PaleoBios. 25(2), 1-7.
Wilson and Allain, 2015. Osteology of Rebbachisaurus garasbae
Lavocat, 1954, a diplodocoid (Dinosauria, Sauropoda) from the early
Late Cretaceous-aged Kem Kem beds of southeastern Morocco. Journal of
Vertebrate Paleontology. 35(4), e1000701.
DOI:10.1080/02724634.2014.1000701
Diplodocinae indet.
(Lapparent and Zbyszewski, 1957)
Tithonian, Late Jurassic
Bombarral Formation, Portugal
Material- (MG 4819; syntype of
Megalosaurus pombali)
posterior mid-distal caudal centrum
(MG 4821; syntype of Megalosaurus
pombali) anterior mid-distal caudal centrum
(MG 4826; syntype of Megalosaurus
pombali) posterior mid-distal caudal centrum
Comments- Lapparent and
Zbyszewski (1957) referred several vertebrae to their new species of Megalosaurus, M. pombali,
including "a vertebra broken in two but entirely of the same type,
although slightly smaller (Pl. XXV, fig. 86), was recovered at São
Gregório de Fanadia (Coll. Geol. Serv.)"
However, Mocho et al. (2016) instead found these did not necessarily
belong to one vertebra and referred them to Diplodocinae indet., as
"the general morphology of these vertebrae is indistinguishable from
that of the middle and posterior caudal vertebrae of the diplodocines Diplodocus, Barosaurus and Tornieria."
References- Lapparent and Zbyszewski, 1957. Les dinosauriens du
Portugal. Mémoires des Services Géologiques du Portugal, nouvelle
série. 2, 1-63.
,
,
and ,
2016.
Systematic
review of Late Jurassic sauropods from the Museu Geológico collections
(Lisboa, Portugal).
Journal of Iberian Geology. 42:, 227-250.
Tornieria
Supersaurus
Dinheirosaurus
Leinkupal Gallina,
Apesteguía, Haluza and Canale, 2014
L. laticauda Gallina, Apesteguía, Haluza and Canale, 2014
Late Berriasian-Valanginian, Early Cretaceous
Bajada Colorada Formation, Neuquen, Argentina
Holotype- (MMCH-Pv 63-1) ~seventh caudal vertebra (85 mm)
Paratypes- (MMCH-Pv 63-2/3) incomplete ?sixth cervical vertebra
(145 mm), incomplete ?eighth cervical vertebra (203 mm)
(MMCH-Pv 63-4) incomplete ?eleventh cervical vertebra (196 mm)
(MMCH-Pv 63-5) partial ?second dorsal vertebra (140 mm)
(MMCH-Pv 63-6) incomplete ~first/second caudal vertebra (60 mm)
(MMCH-Pv 63-7) incomplete ~twelfth caudal vertebra (90 mm)
(MMCH-Pv 63-8) incomplete ~twentieth caudal vertebra (110 mm)
Diagnosis- (after Gallina et al., 2014) proximal caudal
transverse process extremely elongate (about equal or wider to centrum
width) with lateroventral expansions reinforced by robust dorsal and
ventral bars; very robust centroprezygapophyseal lamina in proximal
caudal vertebra; paired pneumatic fossae located on base of
postzygapophysis, opposite to articular side, in proximalmost caudal
vertebra.
Comments- This taxon was found by Gallina et al. (2014) to be a
basal diplodocine in two versions of Whitlock's diplodocoid matrices,
more basal than Tornieria in the former but sister to it in the
latter. Tschopp et al.'s (2015) specimen-level diplodocoid analysis
also recovered it as a diplodocine outside the Galeamopus+Diplodocus+Barosaurus
clade, but with Tornieria, Dinheirosaurus and Supersaurus
more basal.
References- Gallina, Apesteguía, Haluza and Canale, 2014. A
diplodocid sauropod survivor from the Early Cretaceous of South
America. PLoS ONE. 9(5), e97128.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic
analysis and taxonomic revision of Diplodocidae (Dinosauria,
Sauropoda). PeerJ. 3:e857.
Galeamopus Tschopp,
Mateus and Benson, 2015
G. hayi (Holland, 1924) Tschopp, Mateus and Benson, 2015
= Diplodocus hayi Holland, 1924
?= Galeamopus "shellensis" Tschopp, 2013
?= Galeamopus pabsti Tschopp
and Mateus, 2017
Kimmeridgian-Tithonian, Late Jurassic
Morrison Formation, Wyoming, US
Holotype- (HMNS 175; = CM 662; = CMNH 10670) posterior skull,
anterior-posterior cervical vertebrae, cervical ribs,
anterior-posterior dorsal vertebrae, sacrum, proximal-distal caudal
vertebrae, chevrons, scapula, coracoid, sternal plate, humerus, radius,
ulna, carpus, metacarpus, manual phalanges, ilium, pubis, ischium,
tibia, fibula, astragalus, metatarsals
Tithonian, Late Jurassic
Brushy Basin Member of the Morrison Formation, Colorado, US
(Marsh-Felch Quarry 1)
Referred- (USNM 2673; = YPM 1922) incomplete skull, mandible
(Marsh, 1884)
Kimmeridgian, Late Jurassic
Salt Wash Member of the Morrison Formation, Wyoming, US (Bone Cabin
Quarry, Howe Quarry)
(AMNH 969) skull, mandibles, atlas, axis (Holland, 1906)
(SMA 0011; proposed holotype of Galeamopus "shellensis") (young
adult) incomplete skull, mandibles, teeth, hyoid, proatlas, atlantal
intercentrum (25 mm), atlantal neurapophyses, axis (129 mm), third
cervical vertebra (195 mm), fourth cervical vertebra (264 mm), fifth
cervical vertebra (320 mm), sixth cervical vertebra (388 mm), eighth
cervical vertebra (410 mm), ninth cervical vertebra (430 mm), eleventh
cervical vertebra (400 mm), twelfth cervical vertebra (379 mm),
fifteenth cervical vertebra (400 mm), cervical ribs, incomplete first
dorsal vertebra, incomplete second dorsal vertebra (355 mm), partial
third dorsal vertebra, fourth dorsal centrum, partial fifth dorsal
vertebra (157 mm), partial sixth dorsal vertebra, incomplete seventh
dorsal vertebra, incomplete eighth dorsal vertebra, incomplete ninth
dorsal vertebra, incomplete tenth dorsal vertebra, several dorsal ribs,
partial sacrum, incomplete scapula (1.375 m), coracoid, several sternal
ribs, humeri (870 mm), radius (601 mm), incomplete ulna (603 mm),
carpal, metacarpal I (186 mm), phalanx I-1 (53 mm), manual ungual I
(202 mm), metacarpal II (218 mm), phalanx II-1 (78 mm), phalanx II-2
(17 mm), metacarpal III (230 mm), phalanx III-1 (51 mm), metacarpal IV
(208 mm), phalanx IV-1 (35 mm), metacarpal V (180 mm), manual claw
sheath?, partial ilium (885 mm), incomplete pubis (835 mm), partial
ischium, incomplete femur (1.37 m), tibia (845 mm), fibula (850 mm),
astragalus (250 mm trans), metatarsal I (119 mm), phalanx I-1 (69 mm),
pedal ungual I (190 mm), metatarsal II (145 mm), phalanx II-1 (87 mm),
pedal ungual II (157 mm), metatarsal III (158 mm), phalanx III-1 (88
mm), pedal ungual III (144 mm), metatarsal IV (162 mm), phalanx ?IV-1
(36 mm), metatarsal V (148 mm) (Klein and Sander, 2008)
Diagnosis- (after Tschopp et al., 2015) distal end of
paroccipital process curved in lateral view; teeth with paired wear
facets; well-developed anteromedial processes on atlantal
neurapophyses, which are distinct from the posterior wing; atlantal
neural arch bears small subtriangular, laterally projecting spur at its
base; posterior wing of atlantal neurapophyses remains of subequal
width along most of its length; axial prespinal lamina develops
transversely expanded, knob-like tuberosity at anterior end;
interpostzygapophyseal lamina of mid and posterior cervical neural
arches does not project beyond posterior margin of neural arch.
Comments- The holotype was discovered in 1902 and initially
described as a new species of Diplodocus (Holland, 1924). AMNH
969 was initially referred to Diplodocus longus (Marsh, 1884),
USNM 2673 to Diplodocus (Holland, 1906), and SMA 0011 to
Diplodocinae indet. (Klein and Sander, 2008). However, all four of
these (plus Lourinha specimen ML 418) were found to group together by
Tschopp et al. (2015) outside the Diplodocus+Barosaurus
clade, so were included in their new genus Galeamopus. While
SMA 0011 was said to differ from the holotype in several features which
could indicate it is a separate species within the genus, I follow a
differing taxonomic philosophy and would lump all specimens into G.
hayi. Tschopp's (2013) thesis fully describes the specimen and
names it Galeamopus "shellensis", with AMNH 969 as a referred
specimen. While names in theses aren't usually listed in this website,
and this one is only because it was used in the supplementary
information of Tschopp et al.. Tschopp and Mateus (2017)
officially name SMA 0011 Galeamopus
pabsti and instead refer USNM 2673 to it, with AMNH 969 being G. hayi.
References- Marsh, 1884. Principal characters of American
Jurassic dinosaurs. Part VII. On the Diplodocidae, a new family of the
Sauropoda. American Journal of Science (series 3). 27,160-168.
Holland, 1906. The osteology of Diplodocus Marsh. Memoirs of
the Carnegie Museum. 2, 225-264.
Holland, 1924. The skull of Diplodocus. Memoirs of the Carnegie
Museum. 9, 378-403.
McIntosh and Berman, 1975. Description of the palate and lower jaw of
the sauropod dinosaur Diplodocus (Reptilia: Saurischia) with
remarks on the nature of the skull of Apatosaurus. Journal of
Paleontology. 49, 187-199.
McIntosh, 1990. Species determination in sauropod dinosaurs with
tentative suggestions for their classification. In Carpenter and Currie
(eds.). Dinosaur systematics: Perspectives and approaches. Cambridge
University Press. 53-69.
Klein and Sander, 2008. Ontogenetic stages in the long bone histology
of sauropod dinosaurs. Paleobiology. 34, 247-263.
Tschopp, 2013. Evolution of diplodocid sauropod dinosaurs with emphasis
on specimens from Howe Ranch, Wyoming (USA). MS Thesis. Universidade
Nova de Lisboa. 455 pp.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic
analysis and taxonomic revision of Diplodocidae (Dinosauria,
Sauropoda). PeerJ. 3:e857.
Tschopp and Mateus, 2017. Osteology of Galeamopus pabsti
sp. nov. (Sauropoda: Diplodocidae), with implications for neurocentral
closure timing, and the cervico-dorsal transition in diplodocids.
PeerJ. 5:e3179.
Kaatedocus Tschopp and
Mateus, 2012
K. siberi Tschopp and Mateus, 2012
Kimmeridgian, Late Jurassic
Salt Wash Member of the Morrison Formation, Wyoming, US (Howe
quarry)
Holotype- (SMA 0004; "HQ 2") (~14 m, subadult) partial skull,
partial mandibles, proatlas, atlas, axis, third cervical vertebra fused
to cervical ribs (113 mm), fourth cervical vertebra fused to cervical
ribs (131 mm), fifth cervical vertebra fused to cervical ribs (165 mm),
sixth cervical vertebra fused to cervical ribs (194 mm), seventh
cervical vertebra fused to cervical ribs (227 mm), eighth cervical
vertebra fused to cervical ribs (245 mm), ninth cervical vertebra fused
to cervical ribs (270 mm), tenth cervical vertebra fused to cervical
ribs (273 mm), eleventh cervical vertebra fused to cervical ribs (298
mm), twelfth cervical vertebra fused to cervical ribs (314 mm),
thirteenth cervical vertebra fused to cervical ribs (322 mm),
fourteenth cervical vertebra fused to cervical ribs (312 mm)
Referred- (AMNH 7530) skull, mandible, eleven cervical vertebrae
and cervical ribs (Brown, 1935)
(SMA D16-3) partial skull (Schmitt et al., 2013)
Diagnosis- (after Tschopp and Mateus, 2012) U-shaped notch
separating frontals anteriorly (also in Galeamopus); squamosals
restricted to postorbital region; rugose tuberosity marks anterodorsal
corner of lateral surface of posterior cervical vertebrae (also in Suuwassea);
posterior margin of prezygapophyseal articular facet of posterior
cervical vertebrae bordered posteriorly by conspicuous transverse
sulcus, separating facet from prezygapophyseal process.
Comments- The holotype was discovered in 1990-1991 and initially
referred to Diplodocus (Ayer, 2000) or Barosaurus
(Michelis, 2004) before being described as the new taxon Kaatedocus
by Tschopp and Mateus (2012; which is also a chapter of Tschopp's 2013
thesis). AMNH 7530 was among the Howe quarry diplodocids recovered in
the early 1900s, identified as Barosaurus by Brown (1935) and
never described. SMA D16-3 was identified as Kaatedocus based
on braincase morphology by Schmitt et al. (2013). This was verified and
AMNH 7530 was also referred to the genus by Tschopp et al.'s (2015)
specimen-level diplodocid analysis.
Tschopp and Mateus (2012) modified Whitlock's diplodocoid matrix and
found Kaatedocus to be a basal diplodocine sister to Tornieria+Barosaurus+Diplodocus.
In Tschopp et al.'s (2015) superior analysis, it was found to be a
diplodocine sister to Barosaurus.
References- Brown, 1935. Sinclair dinosaur expedition, 1934.
Natural History. 36, 2-15.
Ayer, 2000. The Howe Ranch Dinosaurs. Sauriermuseum Aathal. 96 pp.
Michelis, 2004. Taphonomie des Howe Quarry’s (Morrison- Formation,
Oberer Jura), Bighorn County, Wyoming, USA. PhD thesis. University of
Bonn. 41 pp.
Tschopp and Mateus, 2012. The skull and neck of a new flagellicaudatan
sauropod from the Morrison Formation and its implication for the
evolution and ontogeny of diplodocid dinosaurs. Journal of Systematic
Palaeontology. 11(7), 853-888.
Schmitt, Tschopp, Knoll and Sander, 2013. Paleoneuroanatomy and
braincase morphology indicates the presence of at least two diplodocine
taxa (Dinosauria: Sauropoda) at Howe Ranch (Wyoming, USA). Journal of
Vertebrate Paleontology, Program and Abstracts. 23, 206.
Tschopp, 2013. Evolution of diplodocid sauropod dinosaurs with emphasis
on specimens from Howe Ranch, Wyoming (USA). MS Thesis. Universidade
Nova de Lisboa. 455 pp.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic
analysis and taxonomic revision of Diplodocidae (Dinosauria,
Sauropoda). PeerJ. 3:e857.
Barosaurus
Diplodocus